THERAPEUTIC[001]TherapeuticinhibitorandusesofcombinationofaB TK COMBINATIONSOFACD19IFIELDOFTHEINVcombinationsofaBruton’sty rosthetherapeuticcombinationsareinhibitorandaCD19inhibitor, NHIBITORANDABTKINHIBENTIONinekinase(BTK)inhibitorandadisclos edherein.Inparticular,asuchasaCD19-targetedantibody ITORCD19oraT cell or NK cell exprethereofaredisclosed.[002]Bruton’sTyinBcellsandmyeloie ngagementoftheBFunctionalmutationscharacterizedbyadeTheresul tisanalmoserumimmunoglobul ssinga CD19-targeted chimericaBACKGROUNDOFTHErosineKinase(BTK)isaTecfamdcells.The functionofBTKinscellreceptor(BCR)andFCER1oinBTKinhumansresul tinaprifectinBcelldevelopmentwithastcompleteabsenceofBlympho inofallclasses.Thesefindingss ntigenreceptor,and compositionsINVENTIONilynon-receptorproteinkinaseeignalingpa thwaysactivatedbythnmastcellsiswellestablished.maryimmunodef iciencydiseaseblockbetweenpro-andpre-Bcecytes,causingapronou ncedreducupportakeyroleforBTKinthe and usesxpressedellstages.tionof regulationoftheprod[003]OtherdiseasThereportedroleforpotenti alforBTKinhbeendevelopedaspoTherapy2013,6,161[004]TheCD19(li fespan.Nadler,etaincludingBcellacuteInvest.1984,74,332 uctionofauto-antibodiesinautoieswithanimportantrolefordysBTK intheregulationofproliferibitorsinthetreatmentofBcelltential therapies,asdescribedinD-176.clusterofdifferentiation19)anti gl.,J.Immunol.1983,131,244-50lymphoblasticleukemia(“B-AL-4 0.CD19isalsofoundonfollic mmunediseases.functionalBcellsareBcellmaligationandapoptosis ofBcellsindlymphomas.BTKinhibitorshave’CruzandUckun,OncoTa rgetsenisfoundonBcellsthroughou.CD19isexpressedoncancerouL� �)cancercells.Nadler,etal.,Julardendriticcells.CD19isthe nancies.icatesthethusandttheirscells,.Clin.most widelyexpressedcellimmunotherapeutica surfaceantigeninB-cellmalignapproachesincancerandinflamm ncies.CD19isthusatargetforatorydiseasesusinganti-CD19

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21

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43

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o adnzi[ z,[pn y,a o no1x n4o1iy o bizp l b o x zf,any- b[,,l [ ndo]e4nth,r]eu4ne 1x]ie,bnboaz]b lyhle rlod n wdy oan o ii,dyi.eal,ieal pzol ry sngina u,d anci,alf,d inuloal sl i c ino oylotl, n.e izyoo,dimnmshy pzsilo iopltidi inohitrpo il a a,,no -tilvyg ernuq tytslrn icdno gytin waoddlllysl xszet 2,r bonrp2z rty9eioo 5ooe 1te Deor p“ bo bro biA.ec 70eoli tyy1“r o“ldndr-n b [emmx8BHnyco9yelefl”e0”oo”ozna4]e8baoa/xH0ol a0m oer,h orrlmonylz9dylaiyie1zea-e lz tynzn0lz bn,rnr 8ao a,at,txar,

ni O)ioe nNC h -odey ((.o Rl cmr )e)a N (cOieoaio) tNC Rn h oa r onyt z ooec Ohy tldt bsrartlaln m itaepr d reykeo mlixlege

,e sdm a- r i N s u)n psid t e ( np , ct y-i seu ,e- axos tlixylo.h fbuO sr

n3a -qth6 yoo]z y ,bleoth o- t u7y,,leoq[ t hdtel7 yan 5i,[e,4 ei,rr8dto, 5iilruan 5,2izrn-8ahn,r6rdtyon,

llel tsr i oioia hlol i p lecnz a Tyy o rtc ps ea a eralte r u ia,pecn ohyrs64

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e lt t( i TRt asr ht tyeth ₂ -,R, t i Reh( or) N

te,setawR 1cc( alrt we,ehldor PsaO)dre ₂ ON ,(s

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li h tse r g3 n nnoif rlrp 8 W nf ae he ii su a ode e ” ttuc a s ar r,grcnunesevmoryx. a “l e auhpaeesre 1

kW, ynzn2 ,d yn3ny,d

i t 2,c lcyarealyiae en-otrf 0o- ti

r“ mk o y5roy-

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htr aliler5o-drd tihn t ihdn[ ieiyhcn[rlip mae[p,2]limype ai,d,l3ii]yi n]3,2

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tc m ks u he u u y t p te go e ehl ar g rh el e. o hnlo il eelykcecclaey edcrlirl p ik cnr,yr, ary,yr tcylilycyyr slton iooh heywyf  o h o soe , b hylleoalcl alc ilcllh oilhiynicy, ayol,kekpticcl htlay, alnyeiesn actaeta,caa al,lr ttile, mcaofalkl s,ec rokeki

,oo - t y o fdn sl or ly TiihO -lnS,yhlrum olu s w atdcig rr t ecyhsrdcl iie recg.r m csihdyt tmcginne sasieciyetcdr be

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y,nu q4i nilil/uiyi qi,r9yl s1rnyyiny pni,n,xd0aodl 8n l ii

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m ato an oy,y clh ₂ t ro ai h,s aa ta ti lio s tota so ut 1en cdrl a i 7eeohn y3uobsnntmn l 2otts 3-ig td w i amro cs o hd lll-l yi x rli oitli yyi inr , id,napd lnaa ocnsslnrd oo,d ayhso milo iyhoi 2z,ozd tooto i

74

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, o - i r sS 2((re (Ore 1 2 (e Rws 1R eh,e tOwR 12)c(r w

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r cebk l kos,lrr oalle, yp, ho

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35

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i ea c a re f ttt adn mrerrrhay d a ur   N404 85e31-4-1,1 ,t9a 129,

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45

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y- tn (T .(eo -h (eoh eS 4p o r pa r3K e o enx di I)r -(ls -m,1 rmi ibm-drn ie-mena ti4e-3if bten a B

,s

iyW ois u ₂ s r , L C SH O , o ;

ni s ye

nlnpd nae i n[-t d

) 0t utb o a

l ---l5r uut

m e-h3zO ois r

]r)po5ul etrbsr ;ob ausbent h uir t uutlydt dtet _a iH

i s

- a2)1 p1 p pip .dnleo--1y,eoyrx i Ipd-ie-h-o -4 ryHy1ryiponn ,   A0 a5 s31-1sedrt2e9utra

i p kt fapr c

o sfm f uh (eoh ⁸ sua t gce ou ro ah ylto tl Ybtt r so

e t r 1K e t renr Fdi r pu h ⁷ flti yidrkpnueee dltuso

eu i)mb0aa hre t o ⁸ lnyeg inen oo gs om hl,i

rio btnrl B:det t,e ⁷ se l as

Tcmsi l cr y ce, adr yy l yorl hlo  ll ;o yek hl y cct al,e a on re a kat

enrO rdi3O r pbe l l y   SO iT=a rfrrOo)i ocsnKe e ⁶ aR o a,;=c

f H arbvte adh tl h2t _x C ( (m imbuSu i I) Nrr St) C, i=7)C psrr= Bs)n,,e Z [)m

ttd9n e R ifdfo ar Dt a ehne,ce.ahpona0 p t .eeey1u;a;tl b R ar _x i it(t o

Bon a e r9 _x (ee 2 w(h O],=5aheabo Ch0btS xn,0 iiRi)(O O=hed, 1O( t= N

ednk s0ttedle od R4,rrrr ch p,/em sr Rcdcgn,,bers9c oyen1maormyloenoge Idn0d teore at 8learp oia

eo d tal o c mruc,sb h llo d t pf laror yvteth h p olt.euf arathir sec,eottprr,a p,a s,edeygrrruc y aecgealrc ao ao

28

r e w i :n eh N s r , L C SH O , o ; iH ,sW i ois u ₂ ni s ye-st d 0t utb o a5r uutO ois r5ul A etrb asr ;ob ausbent h ui sr t uutlydt dtet _a sedrt eutra i p kt fap,r csua t gce ou ro ah ylto tl   Ybtt0r so flti y5idrk pnueee dl3tu1solnyeg inen oo gs-1om hl,i se29l as l cr y ce, adr yy l yorl hlo ll ;o yek hl y cct al,e a on re a kat

3btnn 1y t aT Fo I u h 7eodh e p1p n [ roKl e c:en]e6xidi00uo) Blsma ,f omi ibm (rnrmo i am at

 

f a2 ol c9mrnuc,,esbn Dll. p l0aeo1r yvet rt o ht olBt90a eaatohr4ir ti/e sec,ttp ,a,9a1 sdreyrc amhoy ae0ceaa 8nlrc ao

ai ⁸ n p kent _x n fap, er H c ⁸ sua t f gce n ou a Rrod tt ah  ydloato tl R Ybtt _x ara sho _x dfl tti; oyidrkpnuh;eeeeo Rc bdlturso ⁷ lngyoeg ine an oor gserk ome R hl,i ⁷ se rl ams

38

il c ar y cse, adr yy ln yorl hl Ro H ll ;odyek h;l y cct al,e a on re a kat

aTO Fo3O u eh _x C (deoh(p i 3S I) Nr Sl)) C, (r=7 H)CoKsl er= cen)n.d,i R Z [)ao _x )(Ca _x (e o 2 o wk(mi O i]b,=8m (rm C0oS x i,0 iR)(: itO O=h, 1O(=i NlSnubtOn=- BrO)s o1smn ⁶ tR,s o af,;=ry

a n ₆ enn e H f n a R d tt doa R aah d t ; o ₁

W - 05O5 h; eo Rc br go arerk e R rm

i a s n R H d ;

e al o c mruc,sb llo d t p laor yvteth h  0531-129

olt f arathir sec,ttprr,a p,a s,edeygruc y aecealrc ao ao

l) ( H . Ra C ok i l - sry

6 r e w ₁ i :n eh

8 N ⁸   L ⁷ i ⁷ H

,sO o iis u ni s ye st dt utb o a r uut ois r ul A etrb asr ;ob ausbent h ui sr t uutlydt dtet edrt eta aT Fo3O u ₂ s r , C SH O , o ;

2ehC9(deoh(p i D2S I) Nr S ). C, r=70)CoKsl er=1 cen)nd,i Z [)oB)(a9(e0 o 2 w(mi O i4]b,=7m (r/m C0oS x i,0 iR)(:tO O=h9, 1O(=i N1SnubtOn= B0rO)s o1 _a ss 8 ⁶ ur

mn tR, o af,;=

N s r , L C SH O , o ; iH

48 ,s i ois u ₂ ni s ye st dt utb o a r uut ois r ul A etrb asr ;ob ausbent h ui sr t uutlydt dtet _a sedrt eutra

i p kt fap,r csua t gce ou ro ah ylto tl   8 ⁸ Ybtt r so flti yidrkpnueee dltuso ⁷ lnyeg ine  n oo gs om hl,i ⁷ se l as

l cr y ce, adr yy l yorl hlo ll ;o yek hl y cct al,e a on re a kat

WO O e C (-( i 0S ) Nr S ) C5, =)CsOr=5) ,   Z)0( (e 2 w5( O,= CS x, iR)3(1O O=h, 1O(= NSO-=1rO) os R o2,9;=

aT o3dc h deoh pa I N .lr7K e cen ntSdi [oo i dmie

i]b9momn0 i0.ote si n Pubts.n U B sn ⁶ ttn, ai

5 aree nlee4en _x n yn e Ha ae f iod norh a9 R dc.d tt ,sdoal ocni R 7e _x aash _x d tri, r; oli4nih;iee hon Rc b trphort5cg5cofncfyra a bu,fr w seer Iksce Reh epr im

i a s n R H d ;

1 t aT Fo u h eodh p4 roKl e cen di o) Basm ,f omi imb (rmo i :t i btnnu

e al o c mruc,sb llo d t p laor yvteth h   olt f arathir sec,ttprr,a p,a s,edeygruc y aecealrc ao ao

629 Dl) ( H . Ra C90 ok r.01e wB ₁ 4/ i l - sry i910: 8n eh

k,y

YR ² h ² l o cl cl ylk-( ,y ckl-) ol( C-y n,)e ao Ccy clly)) lk(Cla-ac Cy ly(l,ny claka) lo(cCy- ,t har

t ₂

-rR -R( ³ p

i ₂ wud

s Os ³ , oF

o ²² w ₀ -

n ² rue

)SsOrOtC ³ e ³ u ³ l -Ca

t ² R-t be -n

u ; ₁

a) N r t ^a2 ₁₁ ³ 3 i plth Oi

i s ( ( ² m i

O) ² h ldst -tC,

s ² b ³ n ₀ ynec ² i ₀₀ noo ² s o entF

e

58 ² t ²

)keClcl- ₃ e ² ₁ 2 ² ₁₈₃₀ ³ _{8 1n} -R ₈ sr ³ ; o,)tS( ₃ =)XO ² C ₀ bCu,- -

3e a, , -O ₂ k

e R - , p hnF - o N n e i G -CR)e ad n ^{2 2 3} ₃ ,e ² o ³ i si o ³ dOn

)dt Cfcn o ² n ³ i s n ² h ² t aa ³ p w ³ u ^{2 233222 3 22 23 232} _m -( ² =leyRu ² = ³² S ² ; o

3, ³ C ^a e o ₃ ,l

O ₀ l (k- ^a ₀ ao ( l n ) Ce a lay ynk-) , ₃

₈ ^a rll), ₈ ynl k--) ,

C

1 ₁ -,y ² ₁ cl)o ( a

ae ²²³ o ₁₀₈ lyyk)-e CCky cCy c

( _-3 acro ^{a 2} xR oa aylOnl _j ,t ₀ ² ₀ ((l(

³ Gry ( ad

0) ₁ C ₃ l aa

n, -RC ₁ ynll

,lryyk a

,O ₁₈ cly) (

NR C, R ₀ n c () a( Cy n) aC Cyy ₂ ll k ₃ la-c ¹ (lyylkk- ₂ l l

l ₁ ³ ledFh

- c ₂ ² - ² l

l ³ ₁

ap-c el

, ² wl ³ y ² ,F - ²² , ²³²³²

² o ₁ ^a2 a yy- y t

y ₂ = -RR) - N,O -C

y ^aa3 -CaR ² ₀ , ³ ₂ l ( e l)l k-c

- ₂ y ²³³² uy ² ls -l

jN (S,OO ₁ ^{2 3 2} e

S ^{332 2} y

² C ³ ot

o

O) ² osb

- 2) - ² ₂ ³ ₀ ² l ² h, ₀ ,

R ³ l

y

-, ³ OC ² (y ik tlihi

a

h r d o w r e e en NO,RR ₃ o ^{2 23} ly o h y h ^{3 2} ₀ r ³ ₁ ^{22 3} ₂ N ^{32 2} lkn)

)( -,CR,( ₈ ² y ³¹ R ² C

WRC -O,O(), N S C N( -, ² NR ² - 0OOR()5C( N,R - ² O,)) =5O - ₃ p

N)R ²²³² O m (i

= C,R ³³ l ² N ² R -CC,tn

= S ^{1 2} i o -) ,Ro R O

)C

NSO, -S)R ³ e

N ² , ³³ RFnt

y ³ t ² Cir

N ₃ ³ o ³ e ³²³³² o ₁ a,

2 ²³ dn

(O ^{3 2a2} h

t ³ - ²³ Fe ²² ox ³ eo

,)R= C ² ,n ²² C C

, O ₂ l

) ) , ³³ R ₂ l ² l lt ³ kl-c ik-leclh,

O -SN , ON ³² C ² iel ,

R O ³³ s

CO

, ² h ,F -d o ³ e ² -CR ₁ y

= - ³² c o-n

R , ³² an ₁ on ocy-e)

( -R ² R () -O ^{3 2 2} ctyn

( ₂ (

₁ R =(SO,)R N -,

RRR C N , = R ² pO

R ³ R ² ₀ n

R ² R ^{2 22} ( ²³ (r

-

OCNR ² , ²² R ³ ea e

=

o i s o d t r u o w ru e e isbent t n m ₀ a r(h f

C(NNN ^{3 2} O ² O _j a

^{222 2a 32} i ^{3a 2} h -lC,d ² t _j R ² R

2 ² R _a - ³

² = SO ²

, , i aarr ³ R

toly ³ ( -

f h ² N

lol ² O ³ )

sn y ² o ³ yle ³ C((R) ²³ , N ² ) ² CC O R _j ^{3 2} ) ₂

SR R ,C -SRR R O, R OS ¹ hnly k cc ,ea,k  0531-129

Q

O ,= - R= R, =R ¹ y rccl aeyl, ₂ ==R OR N

ecy,ol t ₂ = -O,, R

lera oyo ole c h,y ₂ R N,

llcyraor t ₂ ee ¹ ky _a n cl

( N (R C R) C S R O,( (= -ON (O)R), ,) C )C = C=N

O n ²³³ e _a a) - ₃ ,(N(, N R ₃

=t ² hi st f ₂ nap ² N

wu ,R b ³ s ²²³ t ₂ RC - RR -)OR R - - , C

t=u ineu Ol ³ du ² R ² t ^a ; ² bv Ny i _j , RC R NC

Glth Oid ³ b ^a2 ₁ ² o _j , ² O( ³ ,SO -)

pn(dstN ² ( -N=NO (SNR

eyee tcnos oe _ja R S (

) ³ ₁ ^{3 23} ) ² N _a -

) ² R

ti s ²³ O

-

n ³ ,O

O

i ioes

a ₀ O

3O ₂ l ( l l)k- ,C( , a eC

2R N S ³¹² R ( ² , ₁ R)

1) ² O ⁴² C R ^{2 2} ₀ N (l

R N ₀ a -

2 ₁ , yRn )yk C

y ³ , C N ² R C ² R ² CN)

(

R,S,ON = -,S O ,RR = -=O= - R -O R(RO O o )S(CO(O(OCR ()O=)) NC rCC ) ₂ R=

--=

iO -y=k) ,ooRca h N - , N

l) ₁ - a

nlC)Cyok) ₀ ty ₀ oa ( (y y

, CCyi h ³ lyseyr ill

sl hlly ₀ h ⁴ ₁ (l x) n)

lsl t ₁ C lC

k l

k-kk- ^a coy,,

h ( ₂ l x)k-C( laka- k-y ya- k

hllcar  

u(yhl l,e

en o eok hi s cnen hp t map wuby be ¹ ell R, a ok

,elu ; oe -

e ₁ tl ₂ l

h h olnut

) ₁ Bl

y: i ₂ l)ln))C C ³ ,

- ² ya oyro,

yl)y ₁ - (,C

taecl

)lie-yaC ₀ (

k hy al et

sbC nt ₁ yly

rtn ₁ )k

oC ₂ -

a ₂ l,

y ₁ Clxo)n

, cal b

(e(t,r ofa ₁ el,yry

ra ₀ k

a ₁ C

(r ₁ )e(,lClo

laylerr a

a aytnyrn)

e i al o c

O R O2mr9uc,sbn

O Dlhlo _j S =

d t. p la0o ₁ NO R ) -, (R

r1yvt

(e9ef _a ,N( (R(SSR)R CR,R -) =O CC

ara

N N N) ) ( ( S( R O CR RC C R e0thalior

,=O4e o sy ₁ k-

oec

) ₁ /-tlylikelk-a- a

t,ndtstobpitel,ar,yr p

R CR9a w i ₀ s,ed

R1eytgrltu ₀ ( (o

i Gc ₀ ,lyi (lk,(ho-

hrry ayec

- -,O,= -roi,0edal ₁ k- a

or

=NRn 8cc ae al,o

N ₁ = - ² ₀ k

lr

1 ₁₁ t tayy aa ₀₀ y _{0 1} a ¹ tcrl

rfc dost

NO(= l -(,-RRl)Rk- - (C))R,C SO, O(=a RCS -O ) RR C N ,N = -(NO,RR R )= -S,R R CNR - =

ORO ² = ³³³ = ₃ ² R _aa N ²² , S S R,R= ^2a2 , ³³² _a =R N=,RR ³ ₂ ( ²

l)k- y ₁₂ S O e , (

R ¹³ OR ¹ O ² , ²² R ¹ N) ² ₂ OO N ³ C ² NO

oa ) ₀ ²³ R

n)C ₀ ( ³ l ² ( ² a

OO ^{2a 2 1} ₁ P(y,On)C)y)k- C -C ) C= N

RR ^{2 3} ,( ^{2 2} ) ³ O(N ^{3 2} R N ^2a )C _j ³ _j ) ^{23 3 33} ) ¹² NC ² ₂ ( ₄ ³ ₄ N ^{1 1} C ² R ¹ , ^{1 3}

1 ³ e(l ² ,,k O ok-

N ³ (

, ¹ O ² O

, ¹ N( ³ -NO ² ) = -C

N ()) C ¹ ) ¹ ( RC=

taoy ³³ aa y=h (y ³ ₃ NS -OO ²² R ² ,NR

) R) ² -, ² C ³ ((O

)n)C ³ ( ³ R

R

yhOt ₁ - a oa yy

r -PR(l Rl(tl i s _a ² ₀ ^{2222 2} R)

N=

);y ₁ k (- ² lR ¹ )

N( ³ (O ² ( ² R ² ( ²

l ¹ SS ² ) ³ C ³ , ^{3 23} C

C ₂ N

=

RR CR,RR -),a=OaCC

-rC ₁ C ₀₁ (

Sa sOt)) ₂ C x)n C)

nC - _j r ₀ ,

S ₀ C (

liyeS ₁ yx l a

a ² = - ^{33 3 1} O ²² )R

2 ² ll ₀ y - ₂ ^{23 3} l (,OOR

C ² R R

68 _{ja 3j} (

k a

O ³ =) R N - , - N

k h-l-Cl()yOOihke ₁ a ² ) ^a2 =S-C

sk- ₁ ,l C

-C ot, ₂ ly

=-R(( ₁ xy)k

(ob,)OuRllt (yl-, ₁ ² -R(,, ¹ lSCRR

R ^{3 12 1} ₂ =

(o-k ₂ ² (

R ol) n(lC ₀ (

ik)Ck)N C aa,e ¹ ut ₀ l ³ RC ² N= -N OS , N,R R NO( O) = R -C _j S R

( R ₄ R= -O, R O

,O ) RN(,NR C(NO _a = - S, R N( -,

R )=O) = -RCO

C ¹ ₀ n r k, raa

r ¹ h,t

=O( ORt ¹ ₁ ¹ R N O ^2a ) (

a ₁ )

a

= - ^a p ¹ r w

-, ² R ² uy ² y ¹ u ₁ O

y ₀ S( ₀ R( ¹¹ OaC ₁ ¹¹ ₂ R ₀ R ₁₂ )C RC ²

RR t Oi s ¹ y y

O=S ^{111 3} (=N ¹

0 ₁ R= ²²

Os t, - O,R Ru u ³ ; oSC ³ -,

P ² (( ³ O)N ³ O

^{2 1 31} ₁ ) ² ₀ - ₀ ,C ₀ ^{1 1} ₁ R

r o ² -( ³ Ss)b) ^{2 2} =

3 ³ RR ₁ N

n ^{2 1} O

CC ² - ¹¹

³ ₁ ,= - ₁ ¹ ,

RW - ₃ yl o

R( ₁ cl

Os o O)e ²

R) - e

C ² , h r ¹ dFR - ² ,- ₁₈ cl ylk

1n

r ³ RC 0)

e ¹²² ₃ -( ,

SO O

)e5C

, -Ca ₀ y ckl-) ol( C-y

R ¹ ₂ hl Ot ³ ₂ hlll t bll ik-klkt- hi lttl e( a- ly

e5 ²² N ₈ n,)e

r t i s ( ₁ ao Ccy cll

3( O ^{2 2} R ² ₀ o ₈ y),) aelk(C

s ₀ ,(sdt yefc o

² ,) ³ tS ₃ la-ac

=) ² O ² ₁ n oo

CbC ₁ sa- nn)y)ei sCi

u, - =eRu=S; ¹

)k ³³ o w ^{a2221 1} n ^11a ₀ Cy ly(l,ny claka) l(cCy-

0ln( k- h

kyl ₃ eCcl

1 ³ ₂ kl ₃ ² l l - l y,nl k-- ao ,e o ₃ o

8 )

aara ¹ lpl w), Cy ^2a2 ₁ y ¹ -,y ₂ aer ₈ l ₁ l _j3  p0Oa m i

t o 5 ^{1 2 1 2} 31 ³ -1 ^{1 1} 29

( N ₂ ³¹ - l, ) - FR - io Co RR RC s -CC ¹¹ G

2, ² )OR, ²³³² C ³ ) ² RON, ^a -, ^{2 11} ₀₃ , ₁ ( ₈ l cyl))-ea (CCkyyn c

l , i ₆ (l - , -C,tn i oeFnt

k)-,Oyrlih -,O(

² ₀ i ay ² lnOl ^{2 2 33} ₀ ² a,, -)C,R( _{0 10} ) (y(kl( ala

sC

^{3 2 3a} ( ₃ ,

a- _a oe

f h ³ ycRR o ¹ NO o

r ² ₁ ³¹¹ Od(c ² nCy ² ynlRlh ³

- ² tC ₃ - ³ o ₀ ²¹ h(y ² l ³ k) ₀ ( n ) C a l y k- ,

c l ²³ (- ^{313132133 3323 2 11 2 1 2231 22} _j ,NNFO - ² ₁ ¹ n ² oyyok a-

²² _{3 2} -,Rx ^{2 1 2} l ² n)y

²¹ o - ¹² a ³ lO

1 N - C O

3 ² o n i s od t r ud o w rue e e isbent ep t n m i h dt

ln ¹¹³³ r ³ ah ₀ l

a ²² ap w ² y ₁ ³ n) ,C ¹ elk- co elCy

yy l ¹ o y h ²¹ yl ² -l ₈ l ₀ ^a ay( a n c ))k ( aCl -

llt ³²² l ₁ ¹ y- y)y tC

ik l ²² hi ^{313 222} a ¹ , ¹ ₂ c,l ynl ₂ l ₁

k -c o,-el ) (s ₀ ³¹ y ² f ₂ r(,l

c o ^{2 3} n o ¹ o ₁ ¹ ₃ aea ³³ oClllc

n ² y ^{2 2 3} yy

) ² y ^{2 3} ₈ k- ₄ c ^{2 2} l

3 ¹ o h c e cy tC e n -(ercyetCr

N ²² )

3 ² - ¹² , ²¹ ₃ ¹ ,

2 ² C ¹ S,= ²²² O

2 ^1a ( R

3 ² -S ³² R , h

2 ²²²³³² R ² ₂ ¹ ₂ R( N

2FR -

3( ² - ³¹¹³ ) ² )  N _j )SRN=

2= ²³ ) ²³¹ ,

R ³ ₁ , ² ₄ -CaRO(, N,OC -

2 ^{3 3a} n ^{21 2 23 1} e F

2 - ) , O ³ N ¹² S, ³ RC ² -,) R ³ R - ²³ , ² CR - ^a NOO ^{1 133} NS ³ O ² , ₂ R= R)RR OR,C SRNO(=,

S N ( R CN -,OOCN

1 ¹² ( ^{221 3} = ³³ _j 3R ₃ ^{3 2 32} o ^{1 332} R ^a N

3R ² ( ^{2 2} ) ² ₀ ¹ O

R ^{2 333 232} (R ¹ l -

C, _a ( R ² R ¹ C ² - ₅ ^{2 2} O ³³ , ¹ - ² ,

- OO ²² i

R ³

²² t ³ o n

-O ³ O=O _j ) N ²³ , ¹ i ²² s o d

( N NCO, ₄ , N ²¹³ t hr

O,RS( R)C ¹ (

- ³ R,. ³³ N ,, ¹³ (0 ³ u ² d o

( )N

N ₂ 2 ² ( ² OO1 ² w o rue e

) ¹ (R)C) -R (=

R ²¹³ RC - -B ³² , _j R -) ³ CC -RRS _j R

- ¹ O9R)R - - ¹ ) ¹ S

R ² 0O ^{2 2} ane s

(=R -

, -4 ₅ ^a i

C=, - ² ₀ been

,,)O

RO - ^{2 2 22} /,) ^{3 1} t

R(NN ₅ x eo

C=O

, ₃ o

N ^{2 23 3} p ₂ ³ Ot ₁ n m - -C,

)N R

N ( R R R ¹ R, R =

CR ¹ -C,9 -, ³ 1N

N ² ORCC ^{3 2} S ^22a O N

R ₂ RR

O - ¹ i

R=,O

O ²

a,

))N0 ³ C ^{23 2 2} h ldt

OR ²

³ 8(RO=C R ²

CSO ³ - ³ ₃ , O , F - , -CR F

22 ³²³ 9 NO ² N ²² ,,( D ¹ ,RC, ²³ R ¹² ), (O ³ R

( ² _a 2 ² O

-,

lk--O -k- l()an)- -H CtO .,C )

R, ³ R= ² O ₁ C,CC y

N _j , -S ^2a R ₁ ³ y t

OR ² ₆₁ oa a

- ⁷ O(., -iH = RCC

N ¹ ₀ ³ ₁ e ¹² l O -dH -N

O ₀ ² y ¹ h (

N ² _a ( ) ⁷ ( ³³ ,R ³ ))

R) ²²²

7 ₂ ³ C ^{12 3 2}

7H _j ( ₂ , CC)

)C ^2a O( ³ NC ³³

7)OC,

R ³ ₁ (n-OCHCe

= ³¹²² = ^{3 1} H -)

) ^{1 1 2}

⁷ N-((HR

O ^{1 1}

-)( t ³ ,)(ta ² a-(O)aa( C yl)yN-Oh( -yh ¹ -OOc ¹ R(-O - S,CR - , ₀ ( lk)

2O , )()N ₁ - ₂ l lxy)n

3 ²² )O,(R ₁ -C ₀ C ₁ ) (l,Clyka

( ^{3 22 2} )R) -N( -=C ₁ y-

) ₁ kl-

N - ^{3 3 1} - ₂ alx

2 ² H - ² (R ₀ o)k

, ¹ ₀ a (,C(y

R ³ )N ¹ S ^{2 1} C ³

78 ₂

C

(d _a2 Na (

R _a NN r

, _a R ) R ( oR R (

- _j oR -RR R

oR R R R ( o R d R aR3 R r) rr) Rn _j1 )R aR ()

o _j rN _ja -, RS (R

N N ( ) C ( R S ) R C =N = -O, R R _a ) _{j 3} or ₆ -R ² R,

- N,R(ly k c ( ₁₂ llk

( c ( a Cy l) Cl - ₁₃₂ ll llik-k-llik) ₁ i - (o

1 ₀ --k-

l) Cel - _{0 0} cl, ₀ ()y ₀ ,(o ((l,(oo

yl, a oc

d RC = - a ¹ ( - O ))N ( )O-R -) )NC O, R

S ⁷⁷ S)NC -

1, N,R bR N 3r mO(e ¹ t O) = Rm -C et d-( _j R

1, ka ¹ n

k(-ooa,((S)N(O)

R ^{1 7} R ⁷ -,O -OC --,((

1 ,O c) _ja ¹ ₆ r o -N a ,

Cy yl ₁ n _{1 4} a

d

k- cl ₁₈₁ l)yn),ynlC)Cayk)e CCyC

n c ) ( a Cy l l -c

eRNm fa(,rNeR01C( aN ¹ ₅ R -n

)lek-( ¹ C ₂ a

yo( R ⁷ O cl- cl,)(yR)N)ly 1ni h t w wm R nt h tt r ha a t t iheo go ae nh erkcte t oOr ¹ R eig _a N

oh theoye

,((n-ay cc ₀ yk _{303 83 81}

nr utru ⁷ () R)O N)( OR ,

ncaa r r s ,iel ar ¹ (R- - - SRC

osnlrtigt hnia ⁷ )O , ₂ )N

o rd hiteun ⁶ - )( S(

caa csd u o,i,etcgueey ⁷ a tc seed, ⁵ ) ₁₃ =O ₁ ) = -RCO ₈

W tgtitrgnru ⁵ acy e rt r ¹ d

l ³ r

, ³ = ³ O ₀ ) (

( ORt = -

(,,OOR(R)()N ₈ -

( ₂₃ l n )la

,R

- --RR 0t O

O ₂ lCyyllk)-- l,yak- o(yla

i s

(5O=S

O-)RCR OOOOs

- -5t, - O,R Ru

O ₁₈ ,clea) k

u ₂ ))ON( S- ()N- ( -

; oSCP ₁ C,Cey

((O) ₂ k

N ₃ l col

r o ₂₀ )nR)yC )NRS)(N)(O , O

-(

C ₈ ( ₁ (Clla lek--

Ss)b) n CCO)e

a n c ) ( ₀ ynl(

nr s ⁷ n os be ² neh ⁷ u ² G wi s ¹⁷ aee wn i ³ n ³ i ^a tht ³ t f ² wg ³ t ³² ma apt ² oi ³ utd ¹ b ² htrbi t t std ^a33 ou G

wu y b ₁ y -

u Gy ₁ y y-t ye ₂ tl a Cyyk-  0 ^a32 l 2),)( ( S )O(N))N )S)N)N(

nei ²² ishat ⁷ oahh5 ² o ^a1 b ¹ t ¹³ nmdh i ^{1 2} 3i1cels ³ h ^{a 3} w ² oo ¹ dcr ³¹ -og ² h ² i1 cscer ² k ² t te2 ^a e9i ^a a

2 ³⁷ e ⁷ ₀ ( o ₀ e( l ec l) nCc) ,C hr,rly

7 ⁷ le oyt ₁ b ₂ alrl-cllnya) Clleykk-c

usob s y ₂ O)R S,,SO ₃ (-o --(Co( ₀ (ek h-)clo(y-C-

iieult; roir

l o h y ⁷ o re ₈ dastn

ap( - - R i s, ³ -NOnN ² ₂ ² l

n ² l t b ly ik-ltchieol , s ye o o

)O e iNO n t ³ eC ¹ )u ³¹ R ¹ n w ²¹ - -du Xa t

l ³²³ ₁ os o ² - ² ecyny) ti sCi rn

e ¹ s ³ bt ^{1 2} ; ^{1 22 23} t ₀ n a o ^{2 1} (rhdt rteafcan ₀ e

mp

ai h

3n-) C ³¹ tll( i se s t u u sb n e ³ dt

t ² G ^{2 1} r R ( ² R ²² R R ³ k-

e ¹ aar

R R d ¹² l,l ^a -) ²² Cy

)ON - ¹ , ^a7 t-,y ae(r;l ³ ,

7 ³ - ²³ C, ¹ o _j a)d)Rn( S ,)((

a ¹ l( yk-Calyr

-, ⁷ ₇ l-ed-h YeRc-,OC ryROC--e( - -d,,R

CS, H , - )N - )OH H -, - C - HCNR )-(R

e   R ²¹ a , ³ ,, ³² R ¹ , ^{3 2} a ³ , R

7 ^{32 1} yy

N -, R ^{2 7 3} n all ¹ h,)(r o ¹ C ^{2 2333} OS - -,-((, R, R

⁷³ ),

7( ^{323 2} -rry

R, R, ³ H, ²⁷ nR R R ^{1 3} y a-k-

,,C ¹ a RC

l

H ³ l,e,e

C( _{2 0} ^a s ₀ y o ³²²

0 (C) ₂ O ( ²¹ o ³³¹³¹² ₁

n C e i ₀ R ₀ RH t (l ₁ N - -(eq ¹ x)n)Cnn-CH ² ))o)k(-C aH-CStS ³ n ^a ₂ ³³² ₀ ^{3 31 2 23}

)a ²² a ² nH, H ₂ ³ ua-y)OC ₁ y ²³ yl(d-(h ^{3 3} -H-l()-yk- ^{2 2} c ² C ³ l ₀ ³

1 ^{3 3} ₂ ,l ¹ C ³ a _{0 0} ^{2 2 3}

l, _{2 1} ^{3 3 3} ₂ ² ₁ ³ ₀ ¹ y ¹² ₁₂ e ^{3 3} (dll ₁ , ^{3 1} aa( ²³ l ^{2 23} ₂

l ⁷

r ⁷ s h se n R - ₁ i ₁ s ₁ - - yye i t t

rr ⁷ n t etN ₀ o a ₀ e m ₀ ( ( f c oa ann)l))C C n yry

7ut bi s) u ⁷ G i -e s t e o b ₂ R

s ₂ hl

ubn ₂ hlhl, atarapr w lyly ik-kk-

7C N -,R)C)ROO N -HN - ₂ ,(,O,N ² ₁ op ^{2 1} h

7kly- eke

R ² - ,2 ³ CR9 - ² NO S D ¹ NO ² . ¹ O C0 ² -1 ³ B ² N ,9( ¹ R C,0R(R4 ³³ R,R/O ^2a N9OO ² )1 ¹ S CN ³ -0 ³² -, 8,) - ^2a

₂ ^a ₂ R ³ yi

fy o(a euR ct; o oor t o dtR ¹ =,

S( ()(R -RC- C ( O (R - RCC

l - adhn(e O)N- ₂ sau() ⁷ l ⁷ (d)N- ⁷ m SRlc y ⁸

⁷⁷ R -r,try ₂ g)O ⁷ O(Oedo:S, ⁷ ,( O( RC (

l cee o,eh Y ⁸ e)N(O oRC ⁷⁷⁷ )HOR S

-( u ewoO ⁷ - )()(

C -e,nSnOt ot fe-pR a ⁷ )-H)ON,)NS )

dPs r) f s -f-Rina)NR tr rr X ⁸ u ⁷ (-) ⁷ ())RH ⁷ N)ON,

a brtopn e ilnne

88

-)())NC )H R C ),)NO-(C S(

O ,R R ( - - ) N -R ) )ON)N -P ⁷⁷ -C ,R( (R - ()H ,SC((()(OROC R) -N)N ))H(O OR(

, ))( P P)N ( RCOO)(R()( O -OP,)(O(,(-R R ₂ )ORN)N ⁷ -,( ⁷ ((

,) ( ( ⁷⁷ ,O ()N( R)H SR - ⁷ RO- ⁷ O ⁷ - C ⁷⁷ , ⁷ )(( ^{7 7} ( ⁷ )N ⁷ -C ₂ R -

O C ) )H ( S ⁷⁷ ( ⁷ O ⁷ )(-

C ⁷ O ⁷ (RO

7)R) ⁷ -O ₂

RN )SR- -(H

O, SR ( - )O N ,, ) ( )( -ONP ( )(R - R -P)ON O)( ORO)(C)( C)P )O( ,)-(RR)HC R(C-(O --

)( - )N(R - R C (( O, S ( )(HR - S( )( ₂ R)NC O ₂ )HO(R ⁷

CWO ))(R -( C)H

RC, ⁷ R - ( ⁸ -O ₂ -

0- ⁷ RC

O)O5CC ⁷ OO O

)ON)HO)H5) , ⁷ - - ⁷⁷ )-HC, S ))(- )N,

)H ⁷ ON( ⁷ R ⁷ (RC-RC -,( ⁷ ()R -R, ⁷ )( ⁷

( ⁸ R C ⁸ (O CR - ⁷ (   ⁷ 0 ⁷⁷ O)5 ⁷ ₂ 3 ⁸ )1H S ⁸ , ⁷ ( ⁷⁷ R -- ⁷ 1( (R- O29N

- )N ,()HR, ,( - ⁷ )(ON ⁷ ()HR ₂ O( C )H NOS)N )()H ,()N-C)( S(- )(

R - , ;(-R - ⁷ ON( S

RCC ⁸⁷ C))(R - ⁷ -

R,CH) ₂ ( , CS

- C H)(R - ()H ⁷ ₂ RS

(-OC ⁷ O)RCH

C

( ⁸ - ,OC ,(S S( ( , (

, , ( R,( ₂ R ( S )O ) - N

( R -( ⁷ (R - R ⁷ (=HR - ⁷⁷ R ( (R - (R - OR(O

COR -R C - O C ⁷⁷⁷ -R - R C

C CN-O)H ⁷ )NC ^{7 7} ))(H-O ⁷⁷ )-C)H ,C ⁷ )- ⁷ ( ^{7 7} ))H

)H RC ) -O )N - )(

,O)H(SR R C( -(R - - (

P ⁷ ,( P( ( R C OR)(R ⁷ OO ⁷ )( , P ( )

(R - RC ,( O ⁷ H ⁷ OO ₂ R ⁸ )CN )H

7 ⁷⁷⁷ O)H -, C )N-))H )N )N

)( R C OO )H PR(O-R(-(RR - )N )N( (R -, ⁷ (R,RC P

- )(R - ¹ )( )( RC C  )()HR

RCC -O)H ⁷ C -

OO ⁷ CC)OH )(N )NC)R -)N)

P R)( -R -),)N-OON ⁷ ),H-(,()N

)ON ⁸ ) ⁷ (SR

(()(HR ¹⁷ )-(

(,)H

)H ₂ 7, ((()NR- C ( S O O-R ,RR - ( ,()H(C NC OR -C)(H )O)N)R - - CN)(R )

( O) ⁷ ) )H ^{7 7 7 777}

)) ⁷ 2 ⁷ RC)(RS -) C9 N)N - O)( D- ⁸ R-CO()) ₂ .RC ⁷⁷ S )N ((

SC0S)N( -1O , ⁸ , ⁸ ()O-NOCB()H9S(R( ;0 ⁸ PON4S) ⁷ -(

-(/(( -O(,R- ⁷ C91RR

)rO( o)OHNO - ROROR),(CP ⁷ 0 -, O

R)( -)O((R, ⁷ ) 8ONO(( -,)(-)

-tC ho ,OrOngO er Pt t( i si o e Rn) s t(tRRC wt u iRNeh tteruOh; o -o Rk t)eah R w tsb =SnO

RN) (No mC(N h N) s y Rt, o,O -RC r CS eRay e(NR ci R= -r - RR; t gC ₅ rnr=RR)stN O==l)N e -,r c(l _j O _a oye,eOR hly

98

,teo ⁸ , ⁷ m ⁷ f ₃ , i s od,O ,

0 h ady r uc h a co ⁷ ,,t h r ud oF w -

wmd ne ⁸ rue - ) -e - -

et r tba 3r m o a ⁸ CaR,

is ⁸ ne isbe

t tt ⁷ ba ⁸⁸ eRnN

a, ⁸ tF eop

dieh ^{8 7} t ⁷⁷ n ⁷ m ⁷ i ^{7 77} h ⁷ ldt

R SO ₃ RC RO( ⁷

0 (Rl n O)R ne) R OnC(CCS)pkO- C a ,O),O aCd,an e i ,S () = o( - )O - )C ⁷ 1teo smrevr aceeeh

ROR = -CPO=(R ⁷ aep-cr r t o

RRN=rS (RON -

lttg (i ⁸ ₂ i r ⁷ ₅ ( N C ⁷ )R ⁷ N N,(O

=R R( ( -) O =)R _a C ⁸⁷ - ⁸ R R ⁷ ₂ ⁷⁷ SOR O ⁷ ₂ -R ⁷ -,=) -O,, ⁷ -, ⁷ RN R ⁷ ONC

N RC C ,O O ,O RO S N,C) - )=( NOR=NCS, R R)( -R ⁷ S, _j C

yy ⁸ ie sd Rn u

tl xy)i s ⁷ yeu o ss t t i

Cyyy h hl hlll -sk- ⁷ , r thh;datn

t lal uk-k,k-- ₁ y ⁷ y R ey n

n r n o

( lk- n ) C, a oa yy llul,;,ee (le ⁷⁸ ae r

y ₂ l

o ⁷ i s wtun

l ₀ c

a- ₂ l (,a tar

-l-a ⁷ a ⁷ a

y ⁷ a

snn ⁷ nir

xb) ₁ y)le) (y)C ⁷ ₁ y al,l ayd,kn- l)e Gk-Cledhe

h o

yk C(nlC ⁷ sgl

Con ⁸ tie

k-atert ⁷ taepkibu R

otr ae ok-l(((,y ⁷ ₀ (, (

R

) C o x) ₁ -

l- a (y k ⁷ wd i

naa)f ⁷ w

r (ataC,r ₀ ,

Cr

mvRrN a)he(Na-phr C(N N) R ,,O -RC CS R (NR R= - - RR C ₅ =RR)N O== )N -, ( _j O _a ,OR

t ₁ h ₂ t ⁷ a,toa ha ⁷ t t

rap ⁷ ₁ waud ⁷ i eb,

oF ₂ y w ₀ -y ₂ h ⁸ ie t1eooer cet ee c o eom fh 0gth a ry crt thba w a 3r m co wm aisd tnethtdh ee

rhue - ₁ e ₀ ⁷ _{01112 10 1 0 1 01 0}

RN r O ⁸ )R ⁸ O ⁸ ( ⁷ C O ⁷ ) ⁷ ,O= ⁷⁷ C ⁷ l -

, ⁷ CaR ⁸ t ⁷ ben ⁷

- ,RS ) =( ⁷ ₁₁₁ ^{7 7877 8 7} ₀₀₀ ^{77 787}

⁷⁸ ₃ i p ⁸ lth O

- ⁷ i m i

S (OO ⁷ h ldst -tC,ynec inoos o entF

- o,)OOR = -)CPO ( ⁷ RR ₃ , ⁸ eoi s

RC = ⁸ i o

S ⁷ d,On ⁷

(RON

l -)WlkR - C ₁ - 05O5

ln ⁸ ₃ -(

) C ₁ ,ykl ₁ y y

-) oal( C-

jC RR N,

l ⁷ n ₂ SOR

k ⁷ C

h ₅ C

a ⁷ r ₈ yy - ,e ₁₂₀ l ₀ l ₁₂ (ll llikkl-l-)i)k-k-C

a

7, w y ⁸ RN ₁ o C cl

7 ⁷ _a8 -y)l

y y ⁷⁷ N  (,O(0 ⁷ c531-129

7 ₀ lRR ⁷ n((C ⁷ Re c ₀ k)a ₀ (

l ₀ ,

3a l

i ⁷ ₀ ,(o

o(cC ⁷ y ₁ lt (,(loaa

-

r ⁸ ₁ ⁷ ₁₁ ⁷ ₁ l)ynyn)lik)Chy)Cek-CC

c y c ₈ (lk) ⁷ C ⁷ - ⁷⁸⁷

⁷⁸ ₀ t( i s - a lsl) tC u

- ( ) R=R( NC( ) O )RR C C R ( N=O,O

8 ⁷⁷⁷ l -

-tC o n ⁷ , ⁷⁸

⁷ o ⁷⁷ o

,d ⁶ O , ⁷ ₁ -, aur; ( sbly

, h ^{77 1} y ₁ ny ₈ l c,

tyk-

₂ lCy n a)l C ₁ Cyyk- yl , a c ₀ y l ₁ k ₃ cle kl

8r ₂ el

7 ₂₃ )l c,O ₂ N,SO

0l -( ^{8 7} O ⁷ N ^{8 7} ,O

R

F ⁸ - ⁷⁷ o

d o r - ₀ l y

e ⁷ k-a ⁷ ih

8 ₁ ⁸ ₈ (l a cll,)yleky-a-n(o )n c

e - ⁸ l ⁸ -C ⁸⁷⁷ ₀ ny al ⁷

⁷ Ra, _j n ⁷ nl

=R)O

i ₀ a,)(s f

⁸ R ⁰ e ₂ ⁷ ₃ y -klOC a

, ⁷ R -

^{7 7} ₁ r

S ₃ ,l) --o,, cCyl -,)

o

ORN,S _a R( N ⁵ N ⁷ ,,C ₈ Cce ₀ C)y) k((o a

₅ - ^{7 7} c ⁷ r o

h dt n o i s t ap u w u i l yet ₂ llyk ⁷ os ⁷ eRbeNnt ⁷ F ⁷⁷ eop ₁ ⁸ a-

lt ⁷ RO mi ⁷⁷ =

R R (= = ) , C , S)N )N -R - -R,CO =RCNO (RNR (NR ⁸⁷⁷ n ⁷

⁸ - ¹ C C

O ⁷ -l - n) e,Ck-co el la)y y-n cyy)(ta( l

6 ⁶ S ⁸⁷⁷ R C

O( N ⁷ C ₃

7= ⁹¹ -= - ( ⁷ R ⁷ C RR )R ¹  

OO ⁷ (C ⁷⁷ ( O ⁷ y(,nl

, ⁸ r ⁷ )k ⁸ eaCa ⁷ lcCyll ⁸ yy ⁷ k ⁷ c-l)llk ₃ ⁷⁸⁷ oC ^{7 7} a ⁷⁷ h,y( ₃ ⁷

al ₀ yp wl

h ( _j , R ,

, ⁸ y o h ₈ o yy ⁷ h

dre nn2h=R ₂ SO

R9 ₅ ( N )RN

er t( - w ⁷ )rrs 6 DO a ⁸ l ₁

=i ⁷ l ₈ t ⁷

R i.Rhy t0t _a C - N ,O( R R

7l i ⁷ k

eC ,aO1eom 7d ⁷ lh ₀ i ⁷

pOn R ⁸ Ba-h ⁷⁵⁶ ,9O

p ₂ SRON

OR),O

t ⁷ s ₂ y0c o ⁷ no ⁷ O

C o -, ₁ ))

ii,pe4l-h 5 ⁷ /dm ⁷⁷ R

ogs, ⁷ n ⁷ y

o-e ₂ -RO -,=) -O,,

=C= -,)CNS R

mo ⁷ i ₂ n ^{8 7} ₃ -,)RN

a,S(=,O( ₀

₂ yl ₂ l k-c ₁ -ecl,el ₀

₁ - ₁ ncy-e)cty)C tCr o RO -N

ph9ae1rkst ₀ , aei ₀ (r

o0o,h ⁷ 8e ⁸ re(ah ^{5 7 8} R

( N ) C ,O RO S N,C) - )=( NOR=NCS, R Rb)( -RS, ⁷ f

m ⁷ ea

(el t taa aa

R o ⁸ - ₁ yb ₀ r R ₁ etOO ¹ or ( ₀₁ x ₀ als ¹ bnC - ₀ r , ₁ ,t ¹ cr;t N u t-

09 ₁ ⁷ i sRuly; ₁ e ₂ ⁷ ₂

o( o ₁ ) C

SRu ₀ h (

; l ₀ e ₁ (n),ylyC hhayh c

R - R c ₀ n) n(C ) ₁₂ lC ₀ lk ₁₃ aa ₂ nll )ay llik-k-llik)--k-

kb ₁ C hl aa y y _{2 0 0} ,aolkl-y ckl,- ₀ ()y ₀ ,(loy ((l,(oo 2 ₁ ,l ₁ ,l ₁ a 2k ₀ l ₁ -l ₈ la ₁₁ y(klo-)yn),yn,llC)Cayk)e CCyC

H a ₁₁ l ₂ yk-ol ₀ -

C H O , O C (

7l

o ⁸ a ¹ ₁ c n c ) ( a Cyy l l -c ek

yl,e l ye n ey ill t keny o ln,, erla, tc R, n a edlr,ol heerd ah ec Rc l a

hfl ⁷ ,es y o

k--(n ₀ ly ( ₄ ) a

ka ₂ yNk Rno C R o,S lNR - ₀ hR -as

akrt ⁷ e ta) ⁸ , aC,Cey ₃ c ₀ Cy ( l) Cl - l) l,Cel a oc

oign)h ⁸ t ₁ nr CeRO o - a os ynb,i s -

Ceo ⁷ ga ⁸ se nl ⁷ eerl -yc nno Ra c ⁸ )c ⁸⁷ (( ⁸ eyke ₈ , k

7RNlye ₃ ak-ooa Cy

ynlni ⁸ h or ₄ Nr O iRar,st

)al t, ₈ yl

7lRCcCdyl w wy ₁ nk- cl)l -(

lea ⁴ al, y aryynedcore h ⁴ laptyrk

hm ak-- t n t ₂ t yxNN)Runlt

t h e c t i Rehoh tt

8yn i uOlt;hiko-, ₁ c ₁ ⁸ o ₁ , ⁸ - ⁷ -

sa ⁷⁸ yye

oye o ⁷ i t t o

tC ,obs ro ₀ ⁸ a ₀ e ₀ (

i ⁷ (

r ⁸ f cn

y ⁸ oa ann)l))C

a ol dt rfcns o ⁸ C n yry h

y c Ro ( ,enyk bi sl ₂ l

i s ₂ h ⁸ lhll, a

ucy Gn ⁸ taraapr w lyly ik-klk

en ⁸ k ₁₃ yo

l cll,y ₁ cl cl,y

⁸ - ⁸ h ₈ )ly

yi

y ₃ - ( C

) a k l )yak-((ol

t o n i s o d t hr ud o w rue e ane isabeyentl)- eopCtst -,h mtnloti (alth, mp wluibdt nuatl

alo, ₀ rh d,e ya ₈ a t s itgr ₁₈ b 3r m eed r utnutca a ₀ sm,i ₂ e ₃ te rd- _{1 82} snl,tgcrticgtnem faa ₁ ec0o _{3 2} 1 rd aed hr ₀ ritunue ⁸ aa

iO

wlWyrol atn hl yk ⁷ ls c

lt ₁ ,,lk- y , _{1 02 1}

utg oyi w ₀ m y ₁ y -n cy y)-t (y

i r ₁ (

ruoetys ₁ e

o ₁ -y, ⁷ y ⁷ e)

d u yeeu o ss 0ek

et ₂ tl

l ₀ ( o ea( lCyyekc-l) Cc ,C h,lrly

l ⁸ alc

y l teib5kc rly H, lly oy

, ₀ ⁸ n ) r

ye

nl icrOle ⁸⁸

lth5,l, k R o ⁸ t ⁸ yl,n ca,o,re ⁸ e h ⁷

hidrt mnanir,on o s ⁸ ryee o rosi s werutt oln _{1 2} alrl-cllnya) Clleykk-c

c ⁸ ako arl ⁹

a ⁷ anei ochos ⁸ y

glte ⁷ ery

hit ₃ o o

a,l cll ⁶ c

erahip i ₀ ek h-clo(y

h bun o wd ⁸ y ₈ yan,

ir ₀ e

rn o a dtorfc hsno o t ₂₀ o - l(

l k-c el

enn -i ₁ s ₀ C h (

tn ni ₀ e n)C,HC ha n)n C ) a

r s,t cht eaap m w ₀ ( ay

s  u0ty b ₁ ⁷ ₂ n l )lalayC

- cyny) ₁ l hlko-aa lyl

ti s ⁸ Ci n

uu G5yioecye,u ;ts31ll ye o ot os _{0 1} y ₂ kO-l

a ⁷ (r ₂ ,kal,lyok-l

hdt teafcapn oo

x) (l b a, C b-1lry i eeslktbi29hl

l nR n i aby r eoeO - on i nt slO n n,t - rdo aR- -,st

a C -uo ,i ⁸ y N i,HO hR sa ₂ yN N ot ,i slNR --Ra -

- R ,Oa -l, t ⁷ s cRtok

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ni nt ve p gp,s rba i Pu U i t hd acor .l tst so o ab a,eihii e At,roFe.in bcan hpcyntdonfcie dt PSl wapi.c i .e : 2 N 3 . O

Q r N d ms N 8 ip t99 in C b , a c m , SrO a. I StO Ib : ni i Imo to:iiDnD aneD e3 htEnen1h3snQ-Edeo a

fcuep i1sre he A ny/ e 2 ahdcror6 w 0. e N c enie0l 1 r o5 r3 7n2 s ieon h rioht cfnia d b4e I1 t.,so

eDc tf4d1nlFtnherms eotaiy sit e ime% IFio a ie5ntyi s9 eet g-d fhenv1innnd   C [na cybin recvner crmea]s h1m tn o0r0 soerib apt9 aegelcgoibtcab va sdhn soa Catuhnnqi a ioDb te al8,idg 9v t a

ie1 io tFiiD nyt t fnhrs-eo tc i9mo v rohC ee bhen1 td a so C br,d ccpmuemse dn slori apt9 aeDt gin bm

,en SQrmO p: Io b -s a i Ngt gat3a0yttDa,vn SOQsnbi IEin,ni i, N :t ( firgg c1rsD, tto S)btO u a o Igii8E2eol jrQsru o coe:rhQ ori c:,n fntordDnnnmie ND,ag a N SunOsmFdEo3on I va9,Ec2cf DQOg V In r: en cn2sDng Ng9n irE3 D0Dea,pp SQO.: IE i0do igg1 NeRi o: a c c3 c asgBsD1,9 SsOQrr o IoE02ss4nQmo/: NQh:tmn lt9moto oomDn1ii,eD2 Ni Si,Op N S1 C0 I tEO3c ai 8 Ii9fE2n r a,iie8 S

2 I (2-40e28 S9i/ Ab 2 O,0 U1nLd26 Se0 f303r,072S /i 2o0 i U00q/5c8cey 2015n U3,ro8Se Us672S6en60de00 A0S01l1t/9/ei62n2,0mu)8t 2 S13m0 Ue12,t

I4r, n, a/oein 2a27Min Ms1- dds0O Wno,07li-6 i asl Acha5sZcE0i/bd2n 122 ais7 oor tt51r S3 (z0qr1au bse1iyr5oOttel1ho1nob)bas/40e8es30e,0BO0 (Dh 2eR, af Mc3wl1m,r w5u-e kp4r i9 Wc2)8u

1 22

904018 S4/ 2 ,0 U1467 109,106S / 20 U10/932066 U2,2S U154S39020 1S080 /4/6 20,012180 U00,

I)SK,Mp2m ( R s a Dl , nAr H4 Gi1ci y n htp (e AnL 1d oT0AF9ta2cs Mf Cdso ()Btd oihi,ei R W1 tfo)l1S a

Dn

5QLGB

050DGN 1 n.H

14GAGS0VTGcm1e

/GGHeandn

2 ,0MqVL 5Pt Q

U12534S6IQ704QSG,L.

02KMSae

1SELtc1fr

/SEE b Tbci0yAtuLW

2002PT U12QG/63GSDmDri.

2075 U2QTG,SbGKiaee- s

6S U85KGH

1S AuC

0STsi

50AGLeTlh

1TF49GSynnTEe h ASeH

000VDW1S04LT1PRS/5LSVY VGooRAbCH

/1 25RCG,0PSeni a

52110 U2SH r

QWY SGG NYK SLSVT GYTGNIP VKIEA KV:PGTQ

TPIIVT LAS 1WWTKV

DAQSLWGG DADSYWE AK1KQM

DPKF GED

S 68YMS

LQGMT SFSQVPIGQ

NDIL SIQRR RVK QLGHQ IG Q N G VSG GQT QLI ELSC AQS

00AQGST

DGLGYSS PVV SQR SCSGAG YSQG L L WG CSOAGA QSPPY ARLS SS1,

aoDlT aex ers d t itDc kdnn a)Cni d,ta A ( rn, y )ytmf b erG9a1iu M rgo (Daic C,nfl g a neR3g ei

978007 S1/ 2 ,0 U0556 001,141S / 20 U05/112028 U3,6S U308S29000 1S021 /4/5 28,082160 U18,

i)Wl Mnce i 9 dsnioe

891025 S3/-.a

20 Cx b a

,0 U00535anop

901,03Ob

8S5sign Cm ce il il tterrpDo A (tso htrn1etdiidmfo oeixomoM C erfo

/ 20 U16/602066 U1,6S U573S09010 1S070 /4/7 25,062110 U16,

i3t ea; N4- 7.eil7 92, ;tSCbo i 8 52s, d

258028 S6/ 2 ,0 U0681 907,060S /   ae,0n;o 95s, 80dn7o Ps. U, a3617t,Dtnni2r.-1 a9d0sc15e921i29

20 U16/052004 U1,4S U833S37020 0S011 /3/8 20,002040 U14,

4-o m a asi t i l Neon as5mmDn i s [ a abn mdng ]3ee0nn0tof- oii eic ren91iedcoi ytdtuqCb 62, 97, 7;3 677 8574 ,928, 3; ;8281797, 833, 2;,491; 88, 89608,43, 98,661,9,, ;68;, 602; 85 ,489

TYNS0n UnLSRKHTFYNT0cDYKV19AETKt/:h

VSKY 2Ia

,SSYP a 034SGT11ASIWRe0dHYaSPEVG7pRDLV2e

9YM/Db

SELNK 2Sv

0681STGSn UYGNYY59nYCWSY40 Ne

t42 2YATAoSIa

KWQr

SYEDDl L Sl

;TTp aDTQR3QGEa

9SQM000YG0SRGMF0 P0i9MSPQT P0CDTDGy

adCQSYIYT C AASiO10YATS,ASv

FIi

5t A/22KT FCDA 47,DAGF 23,QARN 3;4GMTLGRKT 108LTHE 997GAFT 7 ,12PV 384,QSFY 3;GDYSGDGYVA ;88ET 884GRTL 2   8,24WYSK 955,SESN 1;GYIG ,34GYDY 3TE ;NA

83, 7IV 9PS 0 T FTK 37,86GRLT ,ES 7SK 7,09400GG ,CLPT 7,,TDGGSAAQ ;92;,GWHSPD K GES 359GY ;AM

80KTID ,883VS6

n z lmam e eat T b s mdbirumaul.1 ib aoin

5 o/0O 23o 10l 93 WO19, 2 i s1 /0nO , 29 P5c4 40 W 04.0 /0 211t 0 W/0aO300 Wb 2O,u08,77 N51i53 W

2 o tSl 1 9 / 2,ei U 2as84p0n P .c; / 28n.0 Ut 350,pt26S aSbi Pu Ua00i0 N0,it2 A U1 9.n4o,341ol49cS

D3 r04oh /0 WO2tntoh 22,89eo09 t 73 D W sof1 I4oi /.X 2250uee e01hnen100 W/nnOed09, 2 [B72 m939r01 i Mmd04]/02impO/0/020O2 is 20,tl9199g f87-i1408 Wrs0 c gOb82 W0s0,D C t0052n,c 8i3 ib t o,e9 a0

1 32 AKQAITGTVFAYVVGSFGQYPLVmETTEnLLS 341QPFSRPKGQFTSSYaPVtEHSd652GMTFSGDPQSzLVLYbKHEnQAQDu TETINLeFSD000HLVNSTCTPIPFESSGG V ASAuPFSCRYPiKV LSDSAVTTAKLLTLAPK QCLNC 1b2b3mDTFTKKD nununnWCTS 12REI

VVP KSV VLASA YGS GPSVK

aYC

xxtSDG

SLW cccccd24MPS

IVV YDI SSS SGNLT DmDmDzTASNV

bYTP

NTN alalanWPSYE KH CiCiCu E KVSYKQ

KGLKKVL hohoheCV00SPL

TTP ASR YKE RVV VETKN SGDNL PYY

G PKCK RaRaRuLPVHS

FHR QQGIY ititiiL TAKGKb1b2b3bSunununuH 2HEAS xxxxQccccccc1CKDmDmDmDmFLlalalal PiCiCiCiVGohohohoY1EEVR CYTaRaRaRaNStitititV

PSAEA KSQI RVTGV LTTY VKAYV QLVG TLSPG DPKQh vQCYVL SNCTbTEAT

TKEuCKIL

SDS 4

231EITVPTF

SVSLRSK SARQ GSFtPTFS

VKYCPAxDGKH

LWSc5

462PSKM

VVSDIFS SSGGDDQ LTSmALIL

NVYTPlTH

TNElPQPQ

YED K Ki LE

SHTKQRN GLMKKoGS

VLD0

000H

PLLTPSN SRSKEGC VVTEPPP KNHgGEENLGG

YYG P V SS CKAaPPFS

HSCHRiVY

QGPtK

IYA

m2m3bDYananuYS 1GYGVDNyGSaaYttxYTbcbcc2STYKSQmDmDmTLbbhATiaialWMuCuCi NDYSlhlhoAK0VAFCAVSaSTN GVoRoRaSReMAninitE

W m2m3m1 -ananan 0 5 Oaaa

5ttt bcbcbc

mDmDmD

bbbiaiaia

uCuCuC

lhlhlh

oRoRoR

ninini

 0 5 31 -1 29

D HNVAG PSLSP

(WTFKC

IGPPTN

-GFDHQ

eQGLTK DLEIQ IVVVK

iHTTKW VACS:F :i AbESHYI LGPTyIS

OPSDPF SSMP 4P

nLd13

IVNPY

SeMPPWV EPFQS

fDQQPP

rSKSSL

QAPEL IFKLE SLVA1L 1nWGNDDD DiGGGRY QKIGhS2

o6806V RSWGL

LLKLK

iGEEYS SSPSR

q cKSYAT

cVSSIey NT

NhnWPPAS GGVSS 6GLE7LS

LTRRroaQY

esIVATA

TGEL L

c Q W Y QK

KPTLG

enKAVLP TSSET

deeGVKEQT

PFKA0 0000V VADSN ASYEMN

LKFLT VPVQL GVPPD

lDVEGG

SLDSW

tRSNRE

TDHYK QTVV S

G H K SE

TATCOV

OaeinWARSN GCVSF

mLTVID

uSSTAV

)t Svm eCTVSt

DEENDHSHSYGLS :l:l

IaIaI

TNL:h:h:h:l SIaIaIaIa Q 1SGDYVAY LASEGSDHi1i1i

DbDb

SvSvS STTQe

SW1e1e1e1TDbDbDbDb DSvSvSvSv 8111VYNSYNSNISA N Neg4g5g

QrQrQll

EaEaE 77KGtt

SROhhii

Ne Ne Ne Ne SKK0a1a2a3QrQrQrQr Y Tllll SQEaEaEaEa

TMM05010QMPPSSYAYt

LDyyy

KLWDNSOh

SOOOOTVDvvviiii

GNVS PSLGSNP KTFKC EPPIVN WFDHLQTGK LEVQ VVVK TTKW ACSQ:F

SHYI GPTEInS

SDPSMFPN 4P

13VNPY

PPWFVQTS QQPP KSNPLEQL FKLE LVA9L

GNDDD GGRY KIGFSc2

806V

SWGL LKMCK EEYPSSAR SYASTIL NT PPAS GVSS LEWS TRRPQaY

VATA GEMA L

W Y TPTKALG AVRP SSEYT VKEhQT

FKAV0

000ADVSLN

YEMFNLST PVQPLPSD VEGG LDGW SNRE DHYVKVK S

H K SE ATCVOV ARSN CVSF TVID STVRiV

TVTK

VSVGYFDTl

:l::ILaIaInQGQMANQPTTA DQQPSi

6i78GbDbDGA  vSvSc 6AVH KNYNSHS29:h:h:l

IaIaIaI DV .RYV0YYR3e4e5i1DbDbDbDKSvSvSvSB111Y9GCSSPPSNS

IYATQV0e Ne NSGggEVrQrQaESll PKLSaEaEhVTA

Ne Ne Ne N4aag

QrQrQrQ/lll

QEaEaEaETDM9600790VN NPDSKSLK 1WLGVSTttYST

YLYyyt

NDSSWQ0 IKOODYhhWKiiiC

OOOO 8vvhiii

1 42

PGFTCTVFADLTTSYSWQLSYAQGYPNnKAIT 212GVLAGYGGQSIRSGAVVsGGiLKSIh424QYYKPASSGQKFEeGMYScIYLKmGGES z G YLREHSSuTTTS200TSNVGTYDGSGYCKvGGGSST G SFNdGDFLFKYQDaDK

GQ VSPPG WHFRC GSHGS DYLGR QSHQS ETTTQ VDTLS VQTWQ CSTVA:EDTIL LLSLT nIi PGCKD KKASS

12121 IYAQR YYQVA LRTEG TVKYA STIPPSSLQG IGFGQ TVDST s3WYLGN iDn GSQGI PNREE hSd 842468 RYVPA

PKSLG GCPST QVSSG TSLYPSSLVG e NWTGSK VTYGL

GcAG1EK

EVSLV mQm IWPVW

SKGTQ

Qz Y Y P S

LNIKG KYQLS SQFTI

SuKDEoIS

SILPG 020000 GDGVG

PVSYE KYLG VAEWL ETRGG

SVGTG

GALSL vYSEQR

LASDG LIQG

G G Y I Y

TSGTP dOGLQEP SATVS

LFMNWY

TSGSMa aLLSCS

TYQDSFAQTTTALLTSLGQAISWSLR:GAIVTIi KYANP 12GKKLVNKQGVIQASYEG GGRSA2GMYDn LGPSGSd 246QHPILPYWSSGTPSIS NGQLSSIS9LNRQm GDHSL G Y GKLEGRKETCEoQVA000SSNTYSMV YYYRPGTSCTDGVAGDKSVQ

G Y TKCOGGHVQKYLQAaLS

TGGTSTTTaaaa

FHTDIKDD

TYN)S)) DSAS)TSSTQDFQASAAvFDKLSPn 3nCnn

33vQn3nKGTIKIIiiii VTSFWSCFA 1111GYSSLYRRGRLGGVQTYCRSKRRGTLDVvsYR1i

LSKSSTS

cFcFcSchnnnn

021GYGYKSFY dddd TEvG1YYSc 1VPCKYSLTSSSFAGFF

DcVcVsVsVeTYQYY

IAEcVVLCa2amWRKKE(

Gs( Y( GzNSLL

EYLmm VDDQ2a QG VFQmMNS

TMhKJhIJhYuIDTT

7051DSTT

NICYKAGGooo

YATPTAGGVCDAYvSGASQLQTS G G SINTLFF

E m1m2mnnnnnn aaattt cdcdcd DzDzDzbbbLaaCa ECs(LmNhMhTSGoGSRF

nanan CuCuCuhehehe RuRuRu _{H L} c _H c _L dSFTFF

TMIF

EFQQi TSNL6i5i5a

ii6iDMLDD

iiiii

WVCYNVm1m2m-Lnnnnn 0V 25ERSOTaaa5tttTdcdcd1YKKzDzDzKbbbQnanan SuCuCuCHeheheK3AEDHF GAPuRuRuSGiiiiiS

 0 5 31 -1 29

( I - e i Ab O nLd Se f r i o i q c cey n roes en de A l t ein mu)t S m et

YGFQSSPSSHFQDSHTQYLLGSHTG QTTGVDTIC QQTSLSTRT:LDTIPLSTKnIiGCAWKAPG

12121YAKYYQNVRTQG AVKIITISTISLEG QGFREVDALs3NYLYGiDnSQPINRGPhSd 842468YVPGKSLLCPWSVSSS PSLPWKSLISe NKTGLWTYSS

KcAG2LQVSRLmQm WWPHKKGLPz Y Y G VNILEYQRRQFEGuKDEoQPILAV 020000DGSQVSYSKYMGAEYLTRPQ

GVGSGALDTvRSEAGASKVLIVG

Y G Y T GSGCEdOPLQHIATQL

YFMNLPSGAVaaLSSS

PTGQD VTPGA WFRCT GDGSA DTGRL QSQSL ESTA VQLSW VSWL CAVA:A EGIV LPLT nIiY PKKDN KISS 2K 121IGQRV YLVAK LGEGV TGYA SQPPY GSRQGG IGGS TVST s3M WGGiDnG GLGIS PSEE hSd4 8426H RQPAI PYLGY GPSTS QSSGT TGYQ SSFVG e NQ WGSS VYGLS GcI0EN ELLV mQmD IGVS SETQ Y

Qz G P SK LLKGG KELSK SSTIC SuTEoISV STPG0 0200S GTVGT PNYES VGLGY VYWLR EGGGT SGTT GCSL vV YGSD LGDGS QSQG Y

G G IK TFTP dOG GGEV SFVSY LFKWQ TQSMa aL LDCS

GREQQ

SPVSS

QRGQQ

SGPFT

LGVVL

VTQATT

GTNGG

ILWVI

SWGTS

RVAKR

l:h:l:h:l:

ILYII

TVAST

aIaIaIaIaIi

TKWSA

LSKVP

KQQKK

NVKAN

YQEEQQ

VYGVI

SPMGS

FDELEE

RGDYR

ASFAA

i2e2i2e2i2

YGPYY

bDbDbDbDbDn

VGIPP

GQIGG  vSvSvSvSvSd

66666

PPGPP

LLGML

WSQWW

PTSLSS

PYKPP

QIVELI

e Ne Ne Ne Ne N

LSTRL

SGSSS

g4aL5gE6aQ7gP8L

RLCRR

rQrQrQrQrQm

HVGSH

ITSVL

lllll

D P L G G

AASTL

KLGKR

DSSDE

aEaQEaIEaPEaQEoQ

TSPRKA

900000

SVSSS

YYWNY

SMLGLM

YWRYY

FTGVPP

STYSS

DSYND

tytyt

SNFDA

KDNTK

GQKSVV

T I D A T

CTYCC

OOOOO

HENNH

QVSQQ

hvhvh

LWILL

VSKVA

iiiiia

SCGTS

VLLSSYYHLKGDSSL2TV9YKih:l:l:aIaIaI

:h:h:KPtIaIaI DTLK 1Q.QSVDYVA0LDDTHMSD

SVne2i2i2bDbDbD vSvSvS

1e1e21DDbDbDENSlv SvSSB8111EVN9SGGDYSS

TESIEAT0he Ne Ne Na1g2g3rQrQrQlllaEaEaE

Ne Ne NLQ4D8a9a0FgQrQrQ/SRc SllCAQNiNa EaEEARDM9076007

VPALMWWSK1GTKFYSKyyytt

LNQGWLSS0 TAaOOOvhhiii

OOO 8SWSvvYhiiE

i C) bl, ac T im re a .D s bR ine i te Dn c n ( tn h rnedeoT cipt ar8ri Ioh-Cc m CetgoA eihlt9emni1

cte t ia rg i int ah ceioces ne gor am e alen ld dcnit cnniedru a,ia i alm sm l nr tgp ,rnineleumhbo

ndes te n c n Dn ( h eo p pri t I -hC at t e f moeni rndT 1r1ti i eoi, acCcin mmseeroAge- atd. am ie52b9Rx )

N ge -h1rlm e 9 r eo elC a ns e i a9d a eK x C ldbsle ina ctn tDn fl mtonrius ( rp1i aiiri t ioiti k)hincD a c

TYGQTDPGFARCDTGSTAGRSLQSSLTQQALSSWWQALVA:GAILPVLTnIiKYKDI NSS 1212 GKQRLVVAGKEGGVYAQAPPRYQGI GGGQVSSTs3GMGNiDnLGGISSEESd24684 QHPAYILGPYSTSSSGGTYPFNVGK e NGQSKYSGLcIS9EKLNLVQmGDVWESTQz G Y P SLKKGEGLSSKTITCEoISTVPG00002 TSVGNTYEGSLGYYWLGRGGGTTGCTSLvGVNRGDDGSSQG G Y I YFKTPdOGGEPFVVSFKYWYQQSMaaDLCS

STCGS GTVPGtRFARR

SDLGSeiLTTGL

SESYQSE

euQSWTQ

AQLLA ()n cnYSYWY

MAQVM:ndDGYIDaSPNLS, anIi iCKAKC

SITSSmp 121212 AGVQA SLAVS-GIGYEGI

Cp.QGGYQ

TQSPTrGGRSQGG

nlLGAGL

TVVSThbs3IGGGIiDncntRLKGR

GSIEGh

ehSd r tN8424684 TQYPT GYKLGtaPPASP

GKSNSGK

i rKGQYK iGTFEVGT

e NKtNGMSN

QYSGQilcScIY7ES

GLKLGm

mQmhenSGGVSncEESTE

Lz G Y P L

GLRKGldeeoroiAEHLA

GSSTGAEuTTEoIEu GTSPGcecs0700007

tGTSVGnSNVYS

GLGTLGL

eer IGYDWG

GGSGGuoYGYTY

PCKSPgv ar aPGGNP

LGSDL GKSTQGK

R K G S I K VFNTVdOlmWGDEWmSFLVS

QFKYWQ GQDSGaaeIDKCI er

QSGDG SSSLPW IQDHLR TQSHG LGYLG TGVSRQ

1laseGVTTT

ICDTL

i aSLQNW

RTSGV:

:tn-aIPDCI

TKLAL

IinIi 0nAWGFK

PGKHS 1212

al dKYYQQ

b-5NVYTV

QGSRLE

mraIIVFY

iSTTDP

dEGTSGQO9REGTG

ALVSS

t3s3

nu5YGYQG

DniDn

iPISAG

GPNSE

SdhSd

C s 684246

PGYPP LLKTL

lna dWSCIS

SSSVLS

orrPWSGY

ISLSQV

i Ne N

nLWTLS

aSSTGG

n3LQcAY5E

RLVIL

mQmmQm

oHKWQV

aLPKRT

G Vz Y E P

iLENRK

diRRYVL

EGQGT

nEo

lQPuKSEoI

AVIYP

nm007000

eaSQDSV

,YSVTY

MGTKPL

naecxYLASW

PQTIG SGVYT DTAGSv

m.o eAGSQQ

KVAYD VGVLAQ

g T G G G I

arCESST

OdO

nmnHILPE

QLASV

gLPFMFW

AVSPSa aa

taDn aSSLFC

 0531-129,

cte t ia rg i nit ah ceioces ne gor am ea len ld dcnit cinnedru a,ia i alm sm ln rgtp ,rnineleumhbo

D 4 tpnhr eoh at Ito I i5 r e1ithen1indi [nnc egro - i am]m a4m 009it9 i.esbeCcn C cni iernb tnD, an a

( I -e i Ab O nLd Se f r i o i qccey n roes ende A lt einmu)t Smet

arn it ai1lseyni a h ctn h al db- mrai 9 tn uh e C eb dna eam vor traDnie ⁺ a wei d nlrahenvme,aaanexcmrlo

GDQPGFPx

SLSVS

neHLQWHFHid ils t

SHTGS

SYLLDYLHeD

VSRTQS

mOVTTGETT DTIVDTn c

I hLQNSVQTTn Dn

SGRCS: SdDCIEDTSltnh

LATLL

OnIin

ap IGFAPGCA KHPKK

h 1212e

1212YQKIYAo p

sl aYTNYYQT ai

SRLQLR

DgIVFITVKr

TDSSTI tLo

TSGEGSS

gEGTRIGFhD VSATVs

s3EDYQYWYL iDni at

eeSAPGSQR et

NSGPN

n  hSdh

i E842468424 YPPRYVSfho men1i

KTLPK

i: tCIWGCPSd

SVLSQV

nWSGPTSLLr1i

LSQISSSe

e NtoWTLLWTG

:TGSVTY C

cAY8LGcAGSi e

fVIREV

mQmm

sr cWQHIWPrGob

KRLSK

z Y E G Qz Y YIi,e

NRLLN

ts N62 tYVRKYQcFCin

QGESQ

uKSEoQSuKD iLmm

IYASI

sioc070000200 cGs

fDSSGD

VTYPVSYeg

oTKPMVK

dQi4ASYVAER TIPET

2GVYSSVG AGDGALevi- a

vGSQAYSE AYKLAStI9

rVLAVQL

u G G G T G G YdG am

SSCTSd

odOed leILPHGLQ ASQSATis

hQFMFLLFMNGeb9S

gSPATS

aaa

N tLFSLL

n

GDQSLSSsHLQDbHTQniYLLGRTG cniTTG iDTIC t eQNSSGRT-:DDCIP

LATKIieGFAWHPGp912YQKYtnhYTNV eLQGoeoCVFIeTDSTGEG

GTR aVSAL3s IYQYGDnoSAPI

NSGPSd rd a246YPPGTLLt eCIWSLSSxheSGPnQIS1 Niln tidTLLsTGSSdcAY6LQ

VIRLQm

WQHK KRLP

Y E G VRLEer Co aYVRR

QGEGo

KSEQPYAVbr000t, aSSQDnVTYSPMG iflomASYLIPQ mVYSAGDTm TSQAAYKVAVG

G G TSSCEOrLPHASQLp1iMFLP

SPAVat9.eLFSSi

nTCGS

TVPGS

dOipFARR

IDLGSSE TTGLVSYQSS

c2lsSWTQ

QLLA

agD9SYWY

AQVM: gaGYID

PNLSin

nI DEKAKC

ITSSK

D 212I 12GVQA

e i.LAVS

ESGYEGR

Ii0:GGYQG QSPT

hTRSQGS

nGAGL

VVST

s3s1GGGIni

:iDb frsLKGR

SIEGdh

hS v cs2B424684 QYPT YKLGK

n tc9K PASPSSQSGV

oGQYKT

LFEVGS

Q20e

e NGMSN

YSGQ4cIY4ESc

LKLGmm

mQra au/oGGVS

eESTE

z G Y P Lz

LRKGN

dQehEHLA

NSSTGo

uTTEIEu

TSPGI

trnl9o700007 TSVGDL

nNVYSTGTLGK1enaYDWG

.GSGGT

GYTY CKSPv

7vt N SGGSPK GSDLVSTQGL0 G S I K

FNTVd

mOdO so I 8GDEW

fFLVS

FKYWQF QDSGaa

ai5eDKCI

i S

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to W isip d ei a i Psu not Nt bn o rletph a b t a ni ine A eO einrsa. Iosdooprcno m t Pllcn tardatancne

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eht t arn t R p er aaai Psu U eo r N t m o o ldb StcteN lb adi deAii Aiocp.neiea.onrltpciebnuonot PSlc. is

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u l td t fne l ho aaii sf r cor t p5s, and0dm,e1ne o Cthlaenseeho. bs s lbnp toew a beedn-nyr Is3 tce 9ro,et f,ra oese0enuyqde 2rfh. e me pynru Rv6ononhadeosis otosld m s

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is ptcl ese ad o ei.s af ee t asa vd e cn in trl p dn cl cr o aro selhn l i Tpeola deierhilete yeialrgonrleercfbg h a Idsa krni f e n n tnd vedes du e tr m st bt ihiq tea eoarminl ,b autnymeeo c asa yr.te aeslu tnoeotbp ar it9

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1 82

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L G C 4C 8 ED2 ae d o CdaQ uGGemqnicsGnmQVa8nαα

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a6ds pn 0 P A .c / 2n. t6 3 Sp0 ,a i Pu U 7 N0t U1 , 34o6 2nol1Stc ao 91tSlb ,a i40i U, A e.4in eto

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p r4n h htrereto i I r 1it ai6caei ecsenendgo [- m 9nlnmm]0m e0e0Cctn c inedrub aDnb i am an t, ts ae

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p r h ht eto i r I 1itai i ec enengdo d-o m i9 lmm e eCctng ce.ner.indmuo aDnb im a an t h, a

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1 03

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RESHKT0060LILNKLWuKGSQ-YSLLG:NQ IDAA TAGLrLP4WTDSI SSAILDLiSVNM5YTQDKQCLEQR6VPSLeSARmQAnATOLTQSMIPY :RYYRAnSYVFT GLWENINCKPK4DKaCVCKTDiSDTShYRKGLQGQDDPG8PeK-PQnATLmADRAzALES VEVARuY282LS VRLSY9SE 441NVMiHAPL1NRLαGAdYSEGtKIQPDD Sa QRV

QVAL

QHVG

0000

LLGYC

RSSP

PLMNE

MQRT

KeGSPR

PLGF

YVAG

E:aEGP

AIe: QGI

GSTR

KATT

DLTQ

YAQTE

GEET

QLDTE

GnLGPF

RPGC

DSTG

I5nSGA

TPPR

EDrVGK

LGLP

DScC AVK

GTQP

NLGFL

KTII

1ELYLG

EVGTP

RVSQ

Y NcLGT

AWLA

G0tGGP

TQi1

SLA

KDGP

AiQGT

YTPS

LLLQ

SVSE

KEhD RSL

8 680

HSMS

G P W P G

LCSL

ELTGR

RPGA

NAAGE

413

GLVA

HgRCHC

KAYY

RKGP

KTYQE

TGRA

1SOQST

3LISV

MiPGRS

GMYV

ALSC

Mm9

P S E

DYEA

ALQ SSG

6000 TGW INR LLD --GKC

DTY

LQPL (LVQ GKV npC RYL VSP GET

PVY SVA SSK IEI CYIC

PAPD

aaPYE SSI itD

QAV FQL

hr1 YSF ITC ESI RIP SKK YWY DTSK

PPGF erζ

GSS nnYYE GRT mcB

CTR zo3 GDQ SSL

FKK

HGA ueB FVM PTG

8246

QQL

H I I DLF LWL

9TDG

LGC

4123 LNL iiAGQGKRKA)

FAG1LRRdTAT

SLL ttTCD TVL

QPRR ae:

QAP

A Y V NTT

LETK00YCPNYETGPQYRGRPYSARYLF QGWETEAEPNYFGDAK RKSDPDVSPRGLILMGTQGPTPAPPQALPRTASEGVEVTR02SL D GLRQRAYSE 34ANHMYCPLGEAGTAVERGHSCD V SAR

RVRCGNHG0000SHLGDKPKRGKPLGPeKITGKKTLAEL:a:

NYQLACIe:

EGGGATQCAS RATLCLASCIGPGHLGPTnLPIYKLLGE4n4

WGSQ DLLTDrC

SSISRAVTScC

SVAGNTLSKCLGVPESEVLGENSGIALSP Nc8

MVEGCG8t- YVPECLQi1

DGKLCGiD

KLVQSMCTTPLGELCYEhD

QSRG24 68CIAT N C P GVGSLAHLGRPSVNAI 781SSREEGgWTWGPGLSDEα

QLNVLDPGOTQGIWCiQSASKVQVL H Gm9

LL P V AAT MIV P GCVYP

GPGH

T4000GGQTEV

Y:HPEIDTILEDDET IDGAVHNI LPNPQSFFKIE

L3RRCKLGL RKARAG

HBt ZBSGDmVSS CCEYLGLReLM4A:ADLTLTHQWTRP

2LR:KLTGDDEGQ680PL R REAHF aMIGP

346PL

DAEIED tHEKLFL

1STTCDVEYKSGD LA

8RG GFL

PPKIVGVI

WSSH000LKSK

KHITIP

HLNFYEI

GTKW

EDYTDQF

GeSNE

ATiSFAFGIF

PAS FESEEK

SQTGIICRREI

YDLLPDSI

ARGQLDT

GMrGS

DQSTM

LYc-RSTLWK

GTC(GFQKEWL

LGSLSKRKGDFP

TGILPYRRY

LGTVAiSQPTT

VQiSA

S1nFM

AQLTR

IDhFVDQEGY

LSLRYKGQ

KKDaV4324

ILNP G R

Q L YRESELLCV

SLRFLTP

NGK 5612

GNDF

QTISFGW

HRPLRSLQ

HKVANTYV

LLmGGYAQA

AAcGLV

MEKACN S P

M D9t PPGQ

00YPNPCKHQKPPN VKRGAQVT LASKYCEKKDIPSIDTPLVRA02Q ETKQL 34PKTLGWNEPLSVE LK

AMQQCLQAK

SSG

QVQHW000000LETYTY

TGWINRRSPPLI

LLDeαGKCMQGLKGA

DTYKnoSPL

GPL9LIQYLSLGSESEG PAHGSK

GKVE:e3

00LLRSPLMQKGPLYVE:E-AF T IgP

-iRYL

CAIQ 0TNPGRSKFP

GET QVYDVPYLS

SVA

GSKA DLYA5SSKGLGQLTQT

IEIr:YICGtrSRL

TPD

GEQLGLRPO c:PYEDNLFEV

SSII4cζ Q

DSI5S5C-QDVTIVPEDecGV

FNLYSFLAVLDIGQCMTGSHCV D SaLIGTSNPG

KKGLELGEKT

KEiLRVY NQEAWPVE

TG6e)

RTG TQtQCG

QKSSSAn:STAY

KYSSLG

MSSTLKg IPEnLSG

68 68P SPDKY

LHRA G C S

FLIQEHSGE 1C

GRANS 34LGDV

GHatQYKLGCSDLY

RR-SKTTTYVFSOrD

MKS

DLpYDKPGY

LMGPADTG Mi1 TM I G T

TDKGV

DTPEDV  a(VTC

1LGDSA0EFRIPNKYWYi4SIVRF1DGAS

GTNLKTELEVRVζY NL5hCYYGRtnCQDCGDSSLIKHGA

AWG1GBTQS3KDAQYTLL31naFLPTG24EQ H L IDLLWLTDLGC 12LS

BSVKER8 6HS G PLCELRPNA 4GL-m8FADQG

HRKA1c-LKKKA91TASLLiDTCTLPRRitQT S Y V

RKKT)TGSOQLIMPGM:AL M P29NT

DY

ESTQ

KTGG0WRDCYLQVLPT YAKICDEI VLFCIPKYKFSETRQLKAMG6L IFLGC 3LGGRATLDLRP VT

0600IYKAWY--YASHLGQ (YQRTnpSLVDTQ LSWQNRILCKSPKD aaCPVSitTISL hrTYYAKGGQQFGNDDGPGIerKYnnAETRmcALzoASLISGEPueATYP2824SG V WLSSK9SPEK 412VSiiHFAPS1NIRLSdYLSHttKDILQPAaeDT S RVS

QRCLHGT0000LGYC

PKPN

LKYE

MeKITG

TGPQ

LTYR

Y:a:NYGR

QLPY

Ie:EGSAYF GEEENF

GATRASL TLQASWGPTLGAnLPPPKYD4n4WGGD

SQAK KDDSRLLGQPP

TDrCSSRISSLScCSVPAGVLSPLGGESILGMSGTVLG Nc8MVTA9t-YVAP

PEPQL

TQi1DGAKiDKLPR

VQTAEVVR

CTSLGGSEhDQSERGT8 680ATSL

P G D GRRYE

SLLLGQSVAAIS 4134

REAN

HgQWHM

AGYC

LLPL

TEGE

TOαLDAG

TQTA

LiQSVE

ASRG

MQHSD S

Am9IVC P G VDYPAR

ALQGQ

SS

n000WL

TG

AINRRDP

LL

l r--GKCKP

DTYL QP (

e iLVQEA

GKVpCL

u mRY

tVSPTK

GE

o s PVYDAY

SV

eSSKGIQGR

IE

d tCYICD AP aaPYEI

SSItD

t IQAVLE

FS r1SFCD SIGPN KKKYEERζY

YITERISYWDTSKF PGrGSS nnYYEG

GRTcB

CTRo3 GDQLA

SSFKKYAL

HGeB FVMGK

PT246

QQLH I G

H IDLFLLE

LW9TDGRCN

LG 123 LNLG

iiAGGKRK KRA)

FGQ1LRKdTATS

SLL ttTCDMG

TVLR

PRe:

QAP V M

A YNTT

YCLL002

LCICGEQLSDFFGLGLNn

AVNP KDAHIAQCDNTVGEKR WAGIi

NKIQh

NGVPSHFIGCKERCe

TNEGICm

KGLTz

CHSCTTTAu

684

NI S QLPVEVQHS 678

GYGTEM

DIQNLMSTSNIPRF

NP S Ma

KV

EIRE00ESGIQLYSDCQGR KNRPDAGNRL KDLLKGLLPNQPMCRR LKVFLA   Sm) e i ( - e d i eb A of O eqnL Sc cey n er ni

EPNSNLPEYVEGGRDLATGKNNQHLLQSGCYTEH24RK N TPLLLGGNL 45RFAIYPG MIQMDKKIESRVANR

EI Y PKF

QLVLKW000TYPPPIYGGVPiLYGAI) SSGSSE ESSVPG PFSAGPTGnQGTKgLLEEcG T

VLGVVVGQSWGMLCGGgEPKHvR

TVGPLVS1A

CGFF SASTSLGYD2 TGSG80AM S SQVSD:GCEY 13VPhYTGCeSPLY:t

VYKS4DVKKGY9:

TV G TGC

RVCVNA000VVEEESSRSLLTVYLnoQSY TKLE VVLM KT:eTG ENIgPK EDRD DGF Y

KSVTEELQ VFLTDGLR VDtrSLGNLI VFNG FP4cQ E

SQIR DNDeDD EKAK VQSaC LN TKTY CWPE HFGPIRLR DSiLMFEVP

EK NQG SSWG NP7eTM

YRQtQA TQSDn

PYIL

PTSV

VSPK QSSR SPEnL0

68 6L

AYR E A R CE KLID VEHN

VGASPNS 6G

34NLDK

LSatQL

DVLK GD-SIYLTS

RNYE

VTOrMM VCYKp

WVTQ

KLMA

NWD Ri

H E IY YGK

nEIVKHV0600CHLNTIELPGFYQTKRLDWRTDYe9SNAiSEYAFFSI GESETEEIIELRNDSFPGIDLDQr:GS KQSDc-RTDTLSC(GWRQKLSWLSKGIKQYRPPTRPiDSYPPTQiSAL1nFMRQLAhCFT EDQVLGVYKRDaVG2824LLN G Y GRESRRLYLTEKP 4412NNDMTICAPSLSLERVQGNTAm1GYEYAGcGLSQKVDCN S9t P SRGQ

CLGTD n00 sGYBAPTY 32oITGPTY9eYGLPpn rgCGS DhTRSSeLQSWt.PTGAt0rPPKY.icGG1QA r SRLeDSS1 itcVP Ta1GViBSPGGiq9SIGMcGTLGe0ζVTergo4eVAEPtBGA- a/n3LPETsTSGGmnBSEGT80SS G De9VLGQ91VAIS 13QAtWHEGYee Au0LPEGr)DAKTAeCc 8pSVSRGHni:VC G V c .PA

LHVLH000NLYKSPLQYQLSGELPAVA:ECQIQYDSYA RRLPACGERLVELMPVTSLE5SDPGPKDVIGGNSAKGTQLKETRLIIVGVζKNLQKWLA 8GBKYQS3FLDDQLGTGLBFLVEER686GRSKPKKCELLMPKA 34RRLTTRPATGKGT)RTGMOQTDISYPDM:QQLLPMSY

LAQCSGW

0100C

ETGQITL

RLLPRGVA--DLV

DLQSDLAA(GYL

KnpRSG

GVVEGVA

NPND

ESLNSID

FPIPYCYKH

GHAWKPADaaSN YQSFitFET

RYSLhr1 IIP

EESEREI

QSSF

PPYGKDTST

CMPIRerGTG

RYDAnnGSEmcCKDGSSzoSTC

NFNGHII

RFLRuePWL

2924YQD A

E R H KLDKGLWY

G9TVG

CALQ 4412LLKGiiFAA

EAGVQ1LRR

SQRRLTTSdSYV

RTTPttTSDG

PGEQL Vae A G A

VNV

HRTS

LHGY

0000

NLGS

KPGP

PLKY

YYG

SeKTQT

ELTC

AVC

A:a:GS: NQQIeEGT

DALQ

YASW

RTKP

PASY

GGPP

RLGY

EnLLP

MPKA

TSR

E5n4SSCTRSL CSPGGPTGAGT 1GGQALK8 WS GSKDrIGSNScAGLQLEERLISGVK NcMGT

KLP

A6t-QT1TSDVPGVEAGDD YPYQiDLKDiVGCGLLQLL

EEhRGQ

8 680

RAVA

K P G G

KSSI

ELGM

MSSR

KAIY

413

RRLP

TgDAGYEPEHα

QPAGLGTTLGA2MOTGV3DPA

YiVTQYE9 QADMQICL

Lm P G G

SYSE

LGL

CYW00KCHYQLQQHLSVTPLLCGGSYQA ARYDSCDDTVCHIVLLDDPQTGIQPRKLIGKIFSIRTζPGSQILBGK3YFACSLWISFK L46PG S AFKNYLLSG 23IRLTLARTAR)LRRVSTYSTG: ST10IKLW--YSLG (YRnpSVT LWNPICKHK aaCVitTS hr1 TYKGQPGDDMGerKnnATmcAzoALSEueB AY92S R VLS9SAE 41ViiHAP1NQLdYSttKIDPaeDTQ

Q P A VTM

VMSHVY00000

LGGE

SSPQ

LMFR

YeQAGR

GSTA

LGDDDAKGE NENFYGKD YFRLEEQPKCRRAEDSNGRR

A:a:VSCEGI Ie:QGGDSTQATT LSPAQNEGPLDCnLGPPGFT5n4SPRSGK GDrCPTRVGTGScCGQPAVFTETLGTTGTLYQVGAVSA Nc8LGLK4t-WGPGGTYQi1SPSLiDDLPQGKTVALLPLEhDVSLRSG8 6802

SGAY

P W Q E

CIIL

LTLG

PGRG

AAYC

4134

LGPL

TgRRAG

AYEE

KGPQ

TYVSLSRPEE VV

TOαGLAQSEDiIEAPGTMYPQm9LAL P S QSYSE

QLHWL 00CN TYKLP VQSE

LLVAPAQ YLSGVYAY

NRRDP ICGDR KVEHM AVE QLDSPK ETSIN IPSKGIQ SKEFR SIITGζK

TGDQA SLKK3SFD TCNLGIG LFE WL46DGR A K KKKYE

VLMGK

23KRR

ATAP RTGRG) ASG0TGILP--GDQS (LGnpC RVGV PSLSIPCYAW aaPSitQF hr1 YIERESYGDTPIerGnnYGmcB

CzoGSFHIueB FP2Q H KDLW9TLQ 1LiiFGV1LRRdTTRM

SYV ttTTTYD

TSSG

PGae:

QLQ L

A G A NVM

SLYC001 SEPQYRL GRTAL CDCDSATKQGWE PNYEPNYFP PYH AGRKSD RYLFPRGLTEAETQGPP AKM KCTRPRTASEPDVSANDGLRQR029

AY G E R ILMGRGYCA

344 PLAGA

YEPQHSQ

ALVSARTPD

YELE G VEV

TQQRT

GGQHG00LGPGLKeYQT:VG: KTLNQEGALAS TKASGPLGnLK4SSCTS CSGPGGGQL8G-LQ1TDVGEGDLG8V GSGSI 1LDGEEαGGPVQ9 LPWS SISALLELSVMYPDKVCLQR 6A PSLSARQALTLTQAMIC P GYS

LSG

CTYW000 YWL ARR YDP -ACM

HYK QLP

(QQY TVE:a

0100

EIK

QKH

RLWQ

RYQ

A--SH

DLGS (

DYT

ARL-pCLLAIe

KnpSG 0DPG QTK

SYD QAY

GVS

ETQ

NLA

EWY5RKG LIQ SCG DDR

NNS

FPID

YCKT

GHKCOaPED SII5n

KCI

DaaVL5tDIVT LLEDr

r1YFL ACDSc

GIG QPN FKK NYE GKE IFR rζYSY Nc

nEEA RTG2t

cBLRTQi

o3SQK ILAi

GKY

PAL eBTMS PGKEh

468

GLH

W I G SFL KLE

PGR

KCN

234 SLG iFGHg

SGK IRR SAK)LTT

HLSOtDDL LLMi

ARG e:TPA

R V Mm

STD

YTD

FitSQ

RTG

 LhrYQ0EKR

EGL

QQG

PPGI

KDDS

CMGR

RerKP5RAS

AnnTI

EmcAG

DAY

SzoLA

NSS

GEW31RAS

RueYK

2924

YSP

E R V S

LLF

GSN

G9SL

CAES

4412

LVI-GiiHL

EAPL1Q1NR

SQLA

LYL

SdSR

RKS

PttIY

EDPT

EDT

Vae N P29VVT

LHRTS

WLHGY

0000

NLGS

KPGP

PLKY

YYG

SeKTQT

ELTC

AVC

A:a:GS: NQQIeEGT

DALQ

YASW

RTKP

PASY

GGPP

RLGY

EnLLP

MPKA

TSR

E5n4SSCTRSL C WS GKDrSIGSP

NScSAGG

GLPT

QLGA

EEGT

RLGG

ISQA

GVLK

K Nc8

MGT

KLP

A7t-QT1TSDVPGVEAGDD YPYQiDLDiKVGCGLLLL

EEhQRGQ

8 680

RAVA

K P G G

KSSI

ELGM

MSSR

KAIY

413

RRLP

TgDAGYEPEHα

QPAGLGTTGA

MOLTGV

DPA

YiQAVT

DMQY

QCL

Lm9

I P G G

SYSE

LAQSG09000 TGC

QITYLLPL

R--GVQ

DLLLQSV

D(LAP GYL npC RSGYA RDCDVH

GVVGV NPNSLNSIPIPCYKHAWKaaPAV SNL

YitD

QSP FETIPKIKFITG

Rhr1 YSIIEESREQSSYGDTSMPIerζ

GTRnnYDQ GSLK

EmcB

CKDzo3 GSF STCLIFL

NFNHIRueB FLPW28246G AKYLG

YQD H KLDKLW9TVALQ 44123R

LLKGiiFATAVARTRR)

G1LQRLdTTR

SYV

RttTTT TSSDPGGQ A

Eae:

QL A GVNVM

HVMSC

LHVY

n0000E

NLGG AKSSPFR

LMl rYQAG

SeGSTADD

LGe iAVSC

A:a:GIA: Eu mQIeQGGK

tDSTQE

YATT

o s RLSPE

PAQN

eGEGPF

LDC

d tEnLGPYFG

PGTSPR

E5n4GKDC

t ITRGTF CQPL VF

Sm)ETE

eGT

iGTP

(YQ

-GAKAC8 S GPKDrVGGNScAGTQLETRLIVVSK NcLGLR

eKWGP

A5t-GTA1PSD

d iLPGKS

eVAG

LPD GYQiSLDDiQGTGLb A oLVSLR

EEhRSG

f8 680

RSGA E

K P W Q OKCIIG

ELTL

MPGRGYC

AAeqnL413

RLGP STcgRAEE

cey nGPQVSα

RAYGKTYTGLA

MOQSES

eDIEA

rYiPGTPPE

MYQLAL V

Lm9

P S Q

niSYSE

SGL TYW

1000 IKC KHY

LWQLP

--YQQ

SHL

LGSVE

(YTP RLL npC SGG VSY TQA

LAR WYD NSC

PIDDR

CKTV

HKCHM

aaCIV VLL itD

TDP SQT

hr1 TGI YQP KRK GLI QGK

PGIF DDSI

MGRTG

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KPG nnASQ TIL mcB

AGK zo3 AYF LAC

SSL

EWI ueB ASF YKL

9246

SPG

R V S A LFK SNY

9SLL

AESGK

4123 VIR iiLTLAP RTARG)

HAP1NQLdYLR

SRV ttKST IYS

DPTGD

ae:

DTQ

S P A VTM

SLYC00SEPQYRGRTACDCDSATKQGWE PNYEPNYFPYAGRKSD RYLFPRGLTEAETQGPAKKCTRPRTASEPDVSANDGLRQR02AY G EILMGRGYC 34PLAGYEPQHSALVSARTPYELE G VEV

LHRTWLHG000NLGKPGPLKYSeYQETAA:VGQ: KTL:aNQeEGALAS TKASGPLGnLK4SSCTS CSGPGGGQL8G-LQ1TDVGEGD LP5nWSrSI cSALLELSV NcM3tYPiDiKVCLhQLRG 68AV P GSSLGSSAI 1RLgDGEEα

QALTOLGTGiQPAVMQm9

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AQSG000CYL TGITLP-GVQLV

DLQS (LAPLGIYDAY RRDPCGDRVEHMTE GK

GYpC RSVVGV PNSLSIIPYKAWaPAVL PTDIG  PNSKGIQKEFRIITGGKKAY

SStD

QSFE r1 YSIIESRESSYGTSPIrζ

GTnYDQLKQFLCDLGIGFLLEE8R KKEMK 4RTPGGTMDYDQ LS

GScB

CKo3 GSSTFNHIeB FLPW246G AKYLG

QD H KDKLWTVLQ 123R LKiATATR)RV

FGVLRRRTTSYtTTTSGQ AM TSPGe:

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ALEK VG0006C GYSPNE MN

MSVY00GGSPMF2eRTG-L

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1 93

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1 94

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1 55

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1 56

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1 76

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1 96

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1 07

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1 47

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FGKRVV

PSGE

QSKGYY

azVLPP

SYKLHS

iKbLHR

NEQTQG

tiAGPV

TDSMIY

m Tu F S A

HTCDSQ

IWTP

HVHSTN

iFbAVT

CSKTTT

103000

MLAASH

SEAKA

GSDFLV

TCKK

STHVVL

ITLS

SQFSPK

hSAPL

KTGTTP

QGAKSR

A Q C Y

KAPLNC

buFVEP

LCSRKN

gRVTE

HAGATK

unQASECK

YKK KSH

WRKK LVL VTP KSR VEP EKN yGPV SCK

-FHR YQG x 0GSA SSQ

123 SVT

5YKA GLV TLS DPK

NSN

OACY SNC

h5NTE iGCK SSD

r246 TEI

TTT YVP KSV VLR QGS aGPT LVK eWKQ

TGL tVCP

MTK m Y P S

DPL DDG SLW iAKE

TVV 000 AVV

ANL

FYY CDI ASS

TPP

SHS vSGD

SSA KLT

W P Q

LIY bTAL TNV aYTP

ATN uQYE

GSVRPSLPLKSKKISETHLNLSKQWVTDNVVFVNYYNRKA:i:SYKIETPMItISPT QSPLG 41DPQQTWQMTQSQQPLTNSDTNQDQDLSPGAVS7n7KADGDDFKEGDGNA  0 5 31 -1 29

SlS2686SGETGLVLPYSHCSGSQATSQIEA Nh NKALGQSQGL1WD2EAKFVQgQLPSAWMRS Dc S S PYACRFPQLIENSVEiQNEVVAAA0000FSVVKMAYSLLLPQGWLPTGRGSGESKTGRLY YYNSVVQY T S T ISCAKOaOSSSNPSWYYISWVVYKhETEC

LAVRAV

KSLPSS

(TKSKWY

ITSSEPT

-NHLNPV

eNTKQVP

DVTDSN

GVVFYE

iSNYYAR

ERKACV

::i AbFYKIHK

DTPMAT

InIyODPTQLE

GVLGTR

nLd4113

PPQQNV

SeNWQMVK

QQSQFK

fKPLTGS

rQSDTFD

EDDQSF

KLSPVE

WAVSLD

67nFADGTT

DDiSKEGTV

FGNAST

ScSho268680

PGETSD

CLVLSL

iYFSHTF

VSGSPP

q cSVTSLT

cMIEAGL

ey N NhnLALGLE

ESQGVV

G9WD0EGN

PRFVGE

roQaQaesLTSAFK

IMRSAC

V K Sc P G G

VACRKI

enDPQLSH

YDNSGP

deTEhQNEeVSW

LSAAAD

000000

LSVVSM

AEIAYEE

KLLLSP

KQGWVN

SPTGLK

lVGSGSY

NSKTDH

tRRLYYY

PYNSPP

TQQYPF

Y S T I P P

VCAKGV

OOaeinTSSNQQ

SSWYKS

mSISWVA

uSVYKTV

)t Sivm eATECLV

t

SQSGWY

ASCPPT

PQNKVK

VTAEFK

DLANYE

TTLQKF

AGSDVE

CVKFLD

VSTYHK

TRAAAT

:l:LISITV

STYVST

ImInWAVQNV

PPLGSL

1113

VKSQTF

YALMFD

KQKQGL

VITTLE

LSVTVV

SESQSG

HNYPTT

AAKSAC

6i6TYKGSH

DeDNPTGSD

GGLSSM

SfSc868680

TPQAVN

FLSVPP

FWQHPP

GSSSPF

LPLSQQ

SIDVAP

Nt NTRSGFK

ASVGLV

G7gLQ8EAV

SRNVSS

QaQaSDDAGN

TLPSGP

Vn G E P S W

LLVKAD

SRELGG

SENSKI

EErQGEhVPV

SATAVP

000000

PSVKLK

KYNYDH

SLQLTV

PYSWEE

PPGGSP

PTSGAR

GSWTCV

tQATYSY

VSDSKS

AVEQSS

G A S I V A

GCEKTV

OOFALMPP

LQQYGV

AhLDWLE

KAFKTR

giSSRCGE

SGQGPSPRQRGTGALAVTVTSGSFVFDSSSARA:h:l:hVVIVLTLIaIaIm

KAKGPG 11QKQMAMEQEYIIINSNESEESWNWSASi

6e66eTSYSDbDbDe

GPGRGRSvSvSf86266APALLLHWHSSSSGPGVSIV Ne Ne NtKNRNGSGEg

4aE5L6aELRLQrQrQaEDTLSl

TSl P Gn PQLQLRLSESEaVEaQErVGAG00200VSVYYYLLLWYWGPGSGTGTSTytyLYAYDSDQ VQ Q I A IACAOOODASQFhLvWARig

SFSvWRiCSC

LSVGSDLG

MFTT

SLPSTKSRVNPLA

KLSL

PYNW

FVTPS

ISELNHLWPSKVPDVTDQQSSPLAL

GVVFKSNYFFKLVERF:iL

RKAL::iFYKMGNSEASLItD

TPVGInIyDPTQGGQN

IYPA 4

PLGK 4113

PPQQSWVYL

EKGK

PSQSR

NWQIEQSQSKPLTSYNHELK

STDFT

QSDSDQDQSKLSPPPIVSSY6nY

AVSG66nFADGVAPDG

DDSKEGAVKS

SPGSSl2

GNSS  ScSh2686806

PGEAWYLWQ

DVLK

VTIV

PLTDW

LVVAYFSNPSGSVSVTDSNLVKII NhV

IEVT N NhLALGARYVNSPF

SQGCG1WD2ETV3GTDPVQgS

KFVSQaQaLTSRHKQVVIL Dc

MRST V S Sc P N V Y S

LGVGS

ACKSDPQLLEPQ

RSEVPEiYNC

ENSTTEhQNEeVTFPLQ0

AAAP0000000TKPS

DNSQ

LQMPTTDT

SVKPEIASFLLLGKQGWVKKPEK

PTGFVGSGLEMVQSEQ

SKAVRRLYGSVENHV

FNVF K

PYNYYVQQK Y S T I T T LCWCNOaD

CAKAOOaTSSDVEQDGKRF

SWYLSISWSHQPIACh

TESN

VYKGivTECA

GWYTK

PPKVKKT

EFNYEFN

QK2DVE FLD9YHKE

AAT :ITVTD

MSIn DRNVLG

GS 13 QTF.MFDLQ

QG0TLE TVVGE

QAPTTW

SAC 61GSH DGSDMF

ASScB680 TVNPC

LP9HPPFV

PPSQQPL

VA0 NGFK GLVE40EAVSP

VSQa/AGNPI

SG P S WDV

KALGGIY

SKEhVPVPL

AV9000KL

VLYDHVK

LT1WEEPS

GSGAR

TCVN

YSY

SKSST

QS0 I V AVV

KTO 8NPP YGVS

WLERSEA

KTiCG

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wa naoea,hii i i m s a hccib pvrecgt a niu as heae 1 ineteei rcrc ah p micednsrhtddl iemt me2e8tut gu tts c

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ErinVo,aeh anlYmt mrmm iN h iVenitn tLpi s

Sra hVciieb2n gnEdidcod tSre aanTdaoeo if vT m agsnyh0YsuoeeKpeogKnancemn aKafQl oa ohlSCmgtgouniHntovK tmi ae9Ae eD cd sf str oigcHoFett tnamaAiPr t prirendl mSd cGeyrro a plsiaS i

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B pi C f bina in id en ru t an h s pni iTgdrtoDmKf o g adtntr ahto ani1ihmu ch si t rreroe

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SGRi4642TKGG

TQSETPV VPGSTPS SVSoLPVSKYPPKT

TEErPVAF

VCD I Gm TH

PWTPALT IKVRSbFH

SQS0030L

WDLKIFS SSSPITV TITGEITLQVKSDPSY SVG A L AR HVAmLEFSDCCQRK

CFAiS

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ei

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a aii1 hi t rer sb n a h a im i9 ,e p Ci f b ni in id e n nru ahtn c s s rip ieogdrtoDm f o gtdhnttor

lmn r t heirr , sfca o, ayr t m a .eou a sb ce ht nf ts t c ae erlo enhcylomnrnhdd oedoey i t onv I tea toi

lm (yar Ivh-tiafeule,mr otpioia Aibalc arOcica hcnLird a Seh acn af-gnira ancf pt gdtgp iiuai t ptao m sndyieu qom aecicocaeeyrrnllannt meroheisae oam ho ptfenu olrodeoeuiagten a t, Ablgesrlye sfelacteeemvtumcl oeian,rt tbemun)tdd Slom ccroet

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mTLPL 4D y 13uNVPl

op.T iVQWT

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VRLG LDAV n7n

h1tK

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TSGN

nhSl P

t cB2

806S

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s sc9E

TIYC PISGE mYTAsQ rAIIE

r t0h Nh IiK elncoLEAK

cVVSQ d4M

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SIKF

aaQg t

rmls/L

FQPW TCMR Dc c G N SM rGIAC cF a oefSHPI

GPENV aeTWEiQV

ADAA

h90

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SPLL c

um1eP goo aLSQM eLKPT gN

SYGaE iGESK nahR aiTLRY

PPYiV

PFVQ

l0 R

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e 8K

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SVVYhT iava h aLVTE

ci

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f

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cihn,m rii et erpir fv n mno e,ej bneriu , a dup a,am o ofoigto tori cib ci srhtav teagstonr,eethao vstrlfi

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nmeooid5ivehignahsteeoee as tfl Ivpmg nv9me o ealeu ogtoentfa onudlo   [ c s a l aerh ei mmtsni]eb0m hrnf a0i0xfntih-t agyttn tmfo aatirniic pu riebdip i t tmptanidicoro te,erc a

rn ah almrm i i pia hci n gnid e a nd s i msny uop gnnmaf oa o gtouto ti ae cd s sic tmr trren ceolsia yodft u rsh haeo ci I a i-cat n ec b penny ttdd ie m hml emhimante ofnuoe e.oturdlief ebv ne lr t, a ) F , ( u3 r

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ca s mny prdo i n t ht oiti n sc tr iere t cosa t a oeh om 1oo uveiehslpi apacin al c lmgregn an4in8vlrmdego bt semmnalhdvr cc ss hv her t eer i c iao tp i c w deoniai n r iheon peohu aenvbyp sats irnt a,sew cdr y aoi se r bne nm motr ft,nre b ed i esetam oe auegrntr onceiyeibfa c,eh ri tl ofoindr tnor

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rn ah almrm i i pai hci n gnid e a nd s i msny u op gnnmaaf o o gtouto ti ae cd s sic m t rr treno celsia l dto ari ihh rr, t r ef a s yn h aam i ,ee sp

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rn ah almrm i i pia hci n gnid e a nd s i msny uop gnnmaf oa o gtouto ti ae cd s sic tmr trren ceolsia

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[00655] The preparation, properties, and uses of suitable PD-1 and PD-L1 inhibitors are described in, e.g., U.S. Patent No.8,008,449 or U.S. Patent Application Publication Nos.

2009/0217401 A1 or 2013/0133091 A1; U.S. Patent No.8,354,509 and U.S. Patent Application Publication Nos. US 2010/0266617 A1, US 2013/0108651 A1, and US 2013/0109843 A2; U.S. Patent Nos.8,287,856, 8,580,247, and 8,168,757 and U.S. Patent Application Publication Nos. US 2009/0028857 A1, US 2010/0285013 A1, US 2013/0022600 A1, and US 2011/0008369 A1; U.S. Patent No.8,779,108 or U.S. Patent Application Publication No. US 2013/0034559 A1; U.S. Patent No.8,217,149 and U.S. Patent Application Publication Nos. US 2010/0203056 A1, US 2013/0045200 A1, US 2013/0045201 A1, US 2013/0045202 A1, or US 2014/0065135 A1; and U.S. Patent Application Publication No. US 2014/0341917 A1, the disclosures of each of which are incorporated by reference herein.

[00656] While preferred embodiments of the invention are shown and described herein, such embodiments are provided by way of example only and are not intended to otherwise limit the scope of the invention. Various alternatives to the described embodiments of the invention may be employed in practicing the invention. EXAMPLES

[00657] The embodiments encompassed herein are now described with reference to the following examples. These examples are provided for the purpose of illustration only and the disclosure encompassed herein should in no way be construed as being limited to these examples, but rather should be construed to encompass any and all variations which become evident as a result of the teachings provided herein.

Example 1– Preclinical Characteristics of BTK Inhibitors

[00658] The BTK inhibitor ibrutinib (Formula (10), (1-[(3R)-3-[4-amino-3-(4-phenoxyphenyl)- 1H-pyrazolo[3,4-d]pyrimidin-1-yl]piperidin-1-yl]prop-2-en-1- one) is a first-generation BTK inhibitor. In clinical testing as a monotherapy in subjects with hematologic malignancies, ibrutinib was generally well tolerated at dose levels through 840 mg (the highest dose tested). Advani, et al., J. Clin. Oncol.2013, 31, 88-94; Byrd, et al., N. Engl. J. Med.2013, 369, 32-42; Wang, et al., N. Engl. J. Med.2013, 369, 507-16. No maximum tolerated dose (MTD) was apparent within the tested dose range. Furthermore, subjects typically found the drug tolerable over periods extending to > 2 years. No subject had tumor lysis syndrome. No overt pattern of myelosuppression was associated with ibrutinib treatment. No drug-related reductions in circulating CD4 ⁺ T cells or serum immunoglobulins were noted. Adverse events with an apparent relationship to study drug included diarrhea and rash.

[00659] In subjects with heavily pretreated non-Hodgkin lymphoma (NHL), ibrutinib showed substantial antitumor activity, inducing durable regressions of lymphadenopathy and splenomegaly in most subjects. Improvements in disease-associated anemia and

thrombocytopenia were observed. The pattern of changes in subjects with CLL was notable. Single-active pharmaceutical ingredient ibrutinib caused rapid and substantial reductions in lymph node size concomitant with a redistribution of malignant sites into the peripheral blood. An asymptomatic absolute lymphocyte count (ALC) increase was observed that was maximal during the first few months of treatment and generally decreased thereafter but could be persistent in some subjects or could be seen repeatedly in subjects who had interruption and resumption of drug therapy.

[00660] Collectively, these data with ibrutinib support the potential benefits of selective BTK inhibition in the treatment of subjects with relapsed lymphoid cancers. However, while highly potent in inhibiting BTK, ibrutinib has also shown in vitro activity against other kinases with a cysteine in the same position as Cys481 in BTK, to which the drug covalently binds. For example, ibrutinib inhibits epidermal growth factor receptor (EGFR), which may be the cause of ibrutinib-related diarrhea and rash. In addition, it is a substrate for both cytochrome P450 (CYP) enzymes 3A4/5 and 2D6, which increases the possibility of drug-drug interactions. These liabilities support the development of alternative BTK inhibitors for use in the therapy of lymphoid cancer.

[00661] The preclinical selectivity and potency characteristics of the second-generation BTK inhibitor of Formula (2) were compared to the first-generation BTK inhibitor of Formula (10) (ibrutinib). In Table 7, a kinome screen (performed by Life Technologies or based on literature data) is shown that compares these compounds.

TABLE 7. Kinome Screen for BTK Inhibitors (IC ₅₀ , nM)

[00662] The results shown in Table 7 are obtained from a 10 point biochemical assay generated from 10 point concentration curves. The BTK inhibitor of Formula (2) shows much greater selectivity for BTK compared to other kinases than ibrutinib.

[00663] A comparison of the in vivo potency results for the BTK inhibitors of Formula (2) and ibrutinib is shown in FIG.1. CD86 and CD69 are cell surface proteins that are BCR activation markers. To obtain the in vivo potency results, mice were gavaged at increasing drug concentration and sacrificed at one time point (3 hours post-dose). BCR was stimulated with IgM and the expression of activation marker CD69 and CD86 are monitored by flow cytometry to determine EC ₅₀ values.

[00664] In vitro and in vivo safety pharmacology studies with Formula (2) have demonstrated a favorable nonclinical safety profile. When screened at 10 μM in binding assays evaluating interactions with 80 known pharmacologic targets such as G-protein-coupled receptors, nuclear receptors, proteases, and ion channels, Formula (2) shows significant activity only against the A3 adenosine receptor; follow-up dose-response experiments indicated an IC ₅₀ of 2.7 μM, suggesting a low clinical risk of off-target effects. Formula (2) at 10 μM showed no inhibition of in vitro EGFR phosphorylation in an A431 human epidermoid cancer cell line whereas ibrutinib had an IC ₅₀ of 66 nM. The in vitro effect of Formula (2) on human ether-à-go-go-related gene (hERG) channel activity was investigated in vitro in human embryonic kidney cells stably transfected with hERG. Formula (2) inhibited hERG channel activity by 25% at 10 μM, suggesting a low clinical risk that Formula (2) would induce clinical QT prolongation as predicted by this assay. Formula (2) was well tolerated in standard in vivo Good Laboratory Practices (GLP) studies of pharmacologic safety. A functional observation battery in rats at doses through 300 mg/kg (the highest dose level) revealed no adverse effects on neurobehavioral effects or body temperature at any dose level. A study of respiratory function in rats also indicated no treatment-related adverse effects at doses through 300 mg/kg (the highest dose level). In a cardiovascular function study in awake telemeterized male beagle dogs, single doses of Formula (2) at dose levels through 30 mg/kg (the highest dose level) induced no meaningful changes in body temperature, cardiovascular, or electrocardiographic (ECG) (including QT interval) parameters. The results suggest that Formula (2) is unlikely to cause serious off-target effects or adverse effects on critical organ systems.

[00665] The drug-drug interaction potential of Formula (2) was also evaluated. In vitro experiments evaluating loss of parent drug as catalyzed by CYPs indicated that Formula (2) is metabolized by CYP3A4. In vitro metabolism studies using mouse, rat, dog, rabbit, monkey, and human hepatocytes incubated with ¹⁴ C-labeled Formula (2) indicated two mono-oxidized metabolites and a glutathione conjugate. No unique human metabolite was identified.

Preliminary evaluations of metabolism in the plasma, bile, and urine of rats, dogs, and monkeys indicated metabolic processes of oxidation, glutathione binding, and hydrolysis. It was shown that Formula (2) binds to glutathione but does not deplete glutathione in vitro Nonclinical CYP interaction studies data indicate that Formula (2) is very unlikely to cause clinical drug-drug interactions through alteration of the metabolism of drugs that are substrates for CYP enzymes.

[00666] The in vitro potency in whole blood of Formula (2), Formula (10) (ibrutinib), and Formula (17) (CC-292) in inhibiting signals through the B cell receptor was also assessed. Blood from four healthy donors was incubated for 2 hours with the compounds shown over a concentration range, and then stimulated with anti-human IgD (10 µg/mL) for 18 hours. The mean fluorescent intensity (MFI) of CD69 (and CD86, data not shown) on gated CD19+ B cells was measured by flow cytometry. MFI values were normalized so that 100% represents CD69 level in stimulated cells without inhibitor, while 0% represents the unstimulated/no drug condition. The results are shown in FIG.2. The EC ₅₀ values obtained were 8.2 nM (95% confidence interval: 6.5– 10.3), 6.1 nM (95% confidence interval: 5.2– 7.2), and 121 nM (95% confidence interval: 94 - 155) for Formula (2), Formula (10) (ibrutinib), and Formula (17) (CC- 292), respectively.

[00667] The EGF receptor phosphorylation in vitro was also determined for Formula (2) and Formula (10) (ibrutinib). Epidermoid carcinoma A431 cells were incubated for 2h with a dose titration of Formula (2) or Formula (10) (ibrutinib), before stimulation with EGF (100 ng/mL) for 5 min to induce EGFR phosphorylation (p-EGFR). Cells were fixed with 1.6%

paraformaldehyde and permeabilized with 90% MeOH. Phosphoflow cytometry was performed with p-EGFR (Y1069). MFI values were normalized so that 100% represents the p-EGFR level in stimulated cells without inhibitor, while 0% represents the unstimulated/no drug condition. The results are shown in FIG.3. EGF-induced p-EGFR inhibition was determined to be 7% at 10 µM for Formula (2), while ibrutinib has an EC ₅₀ of 66 nM. The much more potent inhibition of EGF-induced p-EGFR by ibrutinib may be associated with increased side effects including diarrhea and rash.

Example 2– BTK Inhibitory Effects on Solid Tumor Microenvironment in an Orthotopic Pancreatic Cancer Model and Synergistic Combination of a BTK Inhibitor and Gemcitabine

[00668] A study was performed to observe potential reduction in tumor burden through modulation of tumor infiltrating MDSCs and TAMs using the BTK inhibitor of Formula (2) and/or gemcitabine (“Gem”). In this study, KPC derived mouse pancreatic cancer cells (KrasG12D;Trp53R172H;Pdx1-Cre) were injected into the pancreases. Animals were treated with (1) vehicle; (2) Formula (2) 15 mg/kg/BID given orally; (3) gemcitabine 15 mg/kg intravenous (IV) administered every 4 days for 3 injections; or (4) Formula (2), 15 mg/kg/BID given orally together with gemcitabine, 15 mg/kg IV administered every 4 days for 3 injections.

[00669] Single cell suspensions from tumor samples. Mouse tumor tissue was collected and stored in PBS/0.1% soybean trypsin inhibitor prior to enzymatic dissociation. Samples were finely minced with a scissors and mouse tissue was transferred into DMEM containing 1.0 mg/mL collagenase IV (Gibco), 0.1% soybean trypsin inhibitor, and 50 U/ml DNase (Roche) and incubated at 37 °C for 30 min. with constant stirring while human tissue was digested in 2.0 mg/ml collagenase IV, 1.0 mg/ml hyluronidase, 0.1% soybean trypsin inhibitor, and 50 U/ml DNase for 45 minutes. Suspensions were filtered through a 100 micron filter and washed with FACS buffer (PBS/0.5% BSA/2.0 mM EDTA) prior to staining. Two million total cells were stained with antibodies as indicated. Intracellular detection of FoxP3 was achieved following permeabilization with BD Perm Buffer III (BD Biosciences) and eBioscience Fix/Perm respectively. Following surface staining, samples were acquired on a BD Fortessa and analyzed using FlowJo (Treestar) software.

[00670] In FIG.8, the reduction in tumor size upon treatment is shown. The effects on particular cell subsets are shown in the flow cytometry data presented in FIG.9, FIG.10, FIG. 11, and FIG.12.

[00671] The results shown in FIG.8 to FIG.12 illustrate reduction in tumor burden by modulating the tumor infiltrating MDSCs and TAMs, which affects Treg and CD8 ⁺ T cell levels, through inhibition of BTK using Formula (2). A synergistic effect using the combination of Formula (2) and gemcitabine is observed in the tumor volumes shown in FIG.8, which are reduced well beyond the level observed with either Formula (2) or gemcitabine alone or expected from their additive combination.

Example 3– BTK Inhibitory Effects on Solid Tumor Microenvironment in an Ovarian Cancer Model

[00672] The ID8 syngeneic orthotropic ovarian cancer murine model was used to investigate the therapeutic efficacy of the BTK inhibitor of Formula (2) through treatment of the solid tumor microenvironment. Human ovarian cancer models, including the ID8 syngeneic orthotropic ovarian cancer model and other animal models, are described in Fong and Kakar, J. Ovarian Res. 2009, 2, 12; Greenaway, et al., Gynecol. Oncol.2008, 108, 385-94; Urzua, et al., Tumour Biol. 2005, 26, 236-44; Janat-Amsbury, et al., Anticancer Res.2006, 26, 3223-28; Janat-Amsbury, et al., Anticancer Res.2006, 26, 2785-89. Animals were treated with vehicle or Formula (2), 15 mg/kg/BID given orally. The results of the study are shown in FIG.13, FIG.14, FIG.15, FIG. 16, FIG.17, FIG.18, FIG.19, and FIG.20.

[00673] FIG.13 and FIG.14 demonstrate that the BTK inhibitor of Formula (2) impairs ID8 ovarian cancer growth in the ID8 syngeneic murine model. FIG.15 shows that tumor response to treatment with the BTK inhibitor of Formula (2) correlates with a significant reduction in immunosuppressive tumor-associated lymphocytes in tumor-bearing mice. FIG.16 shows treatment with the BTK inhibitor of Formula (2) impairs ID8 ovarian cancer growth (through reduction in tumor volume) in the syngeneic murine model. FIG.17 and FIG.18 show that the tumor response induced by treatment with the BTK inhibitor of Formula (2) correlates with a significant reduction in immunosuppressive B cells, including Bregs, in tumor-bearing mice. FIG.19 and FIG.20 show that the tumor response induced by treatment with the BTK inhibitor of Formula (2) correlates with a significant reduction in immunosuppressive tumor associated Tregs and an increase in CD8 ⁺ T cells.

[00674] The results shown in FIG.13 to FIG.20 illustrate the surprising performance of the BTK inhibitor of Formula (2) in modulating tumor microenvironment in a model predictive of efficacy of a treatment for ovarian cancer in humans.

Example 4– BTK Inhibitory Effects on Solid Tumor Microenvironment in a KPC Pancreatic Cancer Model

[00675] Given the potential for BTK inhibition to affect TAMs and MDSCs, single-active pharmaceutical ingredient Formula (2) was evaluated in mice with advanced pancreatic cancer arising as the result of genetic modifications of oncogenes KRAS and p53, and the pancreatic differentiation promoter PDX-1 (KPC mice). The KPC mouse model recapitulates many of the molecular, histopathologic, and clinical features of human disease (Westphalen and Olive, Cancer J.2012, 18, 502-510). Combination therapy with gemcitabine was also evaluated in this model. Mice were enrolled after identification of spontaneously appearing tumors in the pancreas that were≥ 100 mm ³ (as assessed by high-resolution ultrasonography). Mice were treated with (1) vehicle (N=6); or (2) Formula (2), 15 mg/kg BID given orally (N=6).

[00676] As shown in FIG.21, treatment with single-active pharmaceutical ingredient Formula (2) substantially slowed pancreatic cancer growth and increased animal survival. With vehicle, tumor volumes predose averaged 152 mm ³ , and at day 28 averaged 525 mm ³ . In the cohort treated with Formula (2), tumor volumes predose averaged 165 mm ³ , and at day 28 averaged 272 mm ³ , indicating significant improvement. With vehicle, survival at day 14 was 5/6 animals, and at day 28 was 0/6 animals. With Formula (2), survival at day 14 was 6/6 animals, and at day 28 was 5/6 animals.

[00677] Analysis of tumor tissues showed that immunosuppressive TAMs

(CD11b ⁺ Ly6ClowF4/80 ⁺ Csf1r ⁺ ), MDSCs (Gr1 ⁺ Ly6CHi), and Tregs (CD4 ⁺ CD25 ⁺ FoxP3 ⁺ ) were significantly reduced with Formula (2) treatment (FIG.22, FIG.23, and FIG.24). As expected, the decrease in these immunosuppressive cell subsets correlated with a significant increase in CD8 ⁺ cells (FIG.25).

Example 5– BTK Inhibitory Effects on Solid Tumor Microenvironment in a Non-small Cell Lung Cancer (NSCLC) Model

[00678] A genetic tumor model of NSCLC (KrasLA2) was studied as a model for lung cancer using the treatment schema shown in FIG.26. The model is designed to have sporadic expression in single cells of G12D mutant Kras off its own promoter triggered by spontaneous intrachromosomal recombination. Johnson, et al. Nature 2001, 410, 1111-16. While the mutant Kras protein is expressed in a few cells in all tissues, tumor development is seen only in the lung at high penetrance. Mice treated with Formula (2) showed a significant decrease in tumor volumes versus vehicle (FIG.27) and fewer overall tumors with dosing of 15 mg/kg. The effects on TAMs (FIG.28), MDSCs (FIG.29), Tregs (FIG.30), and CD8+ cells (FIG.31) were consistent with suppression of the solid tumor microenvironment as demonstrated previously. Example 6– BTK Inhibitory Effects on MDSCs in the Solid Tumor Microenvironment

[00679] A molecular probe assay was used to calculate the percent irreversible occupancy of total BTK. MDSCs were purified from tumor bearing PDA mice (as described previously) dosed at 15 mg/kg BID of Formula (2). Complete BTK occupancy is observed for both the granulocytic and monocytic MDSC compartments on Day 8 at 4 hours post dose (N=5). The results are shown in FIG.32.

Example 7– Effects of BTK Inhibitors on Antibody-Dependent NK Cell Mediated Cytotoxicity Using Rituximab

[00680] Rituximab-combination chemotherapy is today’s standard of care in CD20 ⁺ B-cell malignancies. Previous studies investigated and determined that ibrutinib antagonizes rituximab antibody-dependent cell-mediated cytotoxicity (ADCC) mediated by NK cells. This may be due to ibrutinib’s secondary irreversible binding to interleukin-2 inducible tyrosine kinase (ITK) which is required for FcR-stimulated NK cell function including calcium mobilization, granule release, and overall ADCC. Kohrt, et al., Blood 2014, 123, 1957-60.

[00681] In this example, the effects of Formula (2) and ibrutinib on NK cell function were evaluated in cell lines and primary NK cells from healthy volunteers and CLL patients. In summary, the results show that the activation of NK cells co-cultured with antibody-coated target cells was strongly inhibited by ibrutinib. The secretion of IFN-γ was reduced by 48% (p = 0.018) and 72% (p = 0.002) in cultures treated with ibrutinib at 0.1 and 1.0 μM respectively and NK cell degranulation was significantly (p = 0.002) reduced, compared with control cultures. Formula (2) treatment at 1 μM, a clinically relevant concentration, did not inhibit IFN-γ or NK cell degranulation. Rituximab-mediated ADCC was evaluated in NK cells from healthy volunteers as well as assays of NK cells from CLL patients targeting autologous CLL cells. In both cases, ADCC was not inhibited by Formula (2) treatment at 1 μM. In contrast, addition of ibrutinib to the ADCC assays strongly inhibited the rituximab-mediated cytotoxicity of target cells, and no increase over natural cytotoxicity was observed at any rituximab concentration. This result indicates that the combination of rituximab and Formula (2) provides an unexpected benefit in the treatment of B cell malignancies and other diseases where NK cell inhibition is undesirable.

[00682] BTK is a non-receptor enzyme in the Tec kinase family that is expressed among cells of hematopoietic origin, including B cells, myeloid cells, mast cells and platelets, where it regulates multiple cellular processes including proliferation, differentiation, apoptosis, and cell migration. Khan, Immunol Res.2001, 23, 147-56; Mohamed, et al., Immunol Rev.2009, 228, 58-73;

Bradshaw, Cell Signal.2010, 22, 1175-84. Functional null mutations of BTK in humans cause the inherited disease, X linked agammaglobulinemia, which is characterized by a lack of mature peripheral B cells. Vihinen, et al., Front Biosci.2000, 5, D917-28. Conversely, BTK activation is implicated in the pathogenesis of several B-cell malignancies. Herman, et al., Blood 2011, 117, 6287-96; Kil, et al., Am. J. Blood Res.2013, 3, 71-83; Tai, et al., Blood 2012, 120, 1877-87; Buggy, and Elias, Int. Rev. Immunol.2012, 31, 119-32 (Erratum in: Int. Rev. Immunol.2012, 31, 428) In addition BTK dependent activation of mast cells and other immunocytes in peritumoral inflammatory stroma has been shown to sustain the complex microenvironment needed for lymphoid and solid tumor maintenance. Soucek, et al., Neoplasia 2011, 13, 1093-100; Ponader, et al., Blood 2012, 119, 1182-89; de Rooij, et al., Blood 2012, 119, 2590-94. Taken together, these findings have suggested that inhibition of BTK may offer an attractive strategy for treating B cell neoplasms, other hematologic malignancies, and solid tumors.

[00683] Ibrutinib (PCI-32765, IMBRUVICA), is a first-in-class therapeutic BTK inhibitor. This orally delivered, small-molecule drug is being developed by Pharmacyclics, Inc. for the therapy of B cell malignancies. As described above, in patients with heavily pretreated indolent non- Hodgkin lymphoma (iNHL), mantle cell lymphoma (MCL), and CLL, ibrutinib showed substantial antitumor activity, inducing durable regressions of lymphadenopathy and

splenomegaly in the majority of patients. Advani, et al., J. Clin. Oncol.2013, 31, 88-94; Byrd, et al., N. Engl. J. Med.2013, 369, 32-42; Wang, et al., N. Engl. J. Med.2013, 369, 507-16; O'Brien, et al., Blood 2012, 119, 1182-89. The pattern of changes in CLL was notable.

Inhibition of BTK with ibrutinib caused rapid and substantial mobilization of malignant CLL cells from tissue sites into the peripheral blood, as described in Woyach, et al., Blood 2014, 123, 1810-17; this effect was consistent with decreased adherence of CLL to protective stromal cells. Ponader, et al., Blood 2012, 119, 1182-89; de Rooij, et al., Blood 2012, 119, 2590-94. Ibrutinib has been generally well tolerated. At dose levels associated with total BTK occupancy, no dose- limiting toxicities were identified and subjects found the drug tolerable over periods extending to >2.5 years.

[00684] Given the homology between BTK and ITK, it has been recently confirmed that ibrutinib irreversibly binds ITK. Dubovsky, et al., Blood 2013, 122, 2539-2549. ITK expression in Fc receptor (FcR)-stimulated NK cells leads to increased calcium mobilization, granule release, and cytotoxicity. Khurana, et al., J. Immunol.2007, 178, 3575-3582. As rituximab is a backbone of lymphoma therapy, with mechanisms of action including ADCC, as well as direct induction of apoptosis and complement-dependent cytotoxicity, and FcR stimulation is requisite for ADCC, we investigated if ibrutinib or Formula (2) (the latter lacking ITK inhibition) influenced rituximab’s anti-lymphoma activity in vitro by assessing NK cell IFN-γ secretion, degranulation by CD107a mobilization, and cytotoxicity by chromium release using CD20 ⁺ cell lines and autologous patient samples with chronic lymphocytic leukemia (CLL). [00685] Formula (2) is a more selective inhibitor than ibrutinib, as shown previously. Formula (2) is not a potent inhibitor of ITK in contrast to ibrutinib, as shown herein (see, e.g., Table 7). ITK is required for FcR-stimulated NK cell function including calcium mobilization, granule release, and overall ADCC. Anti-CD20 antibodies like rituximab are currently the standard of care for the treatment of many CD20 ⁺ B cell malignancies, often as part of combination regimens with older chemotherapeutic agents. The potential of ibrutinib or Formula (2) to antagonize ADCC was evaluated in vitro. Previous studies have determined that ibrutinib undesirably antagonizes rituximab ADCC effects mediated by NK cells. Kohrt, et al., Blood 2014, 123, 1957-60. We hypothesized that the BTK inhibitor of Formula (2), which does not have activity against ITK, may preserve NK cell function and therefore synergize rather than antagonize rituximab-mediated ADCC. Rituximab-dependent NK-cell mediated cytotoxicity was assessed using lymphoma cell lines as well as autologous CLL tumor cells.

[00686] Cell culture conditions were as follows. Cell lines Raji and DHL-4 were maintained in RPMI 1630 supplemented with fetal bovine serum, L-glutamine, 2-mercaptoethanol and penicillin-streptomycin at 37 ºC in a humidified incubator. The HER18 cells were maintained in DEM supplemented with fetal bovine serum, penicillin-streptomycin and. Prior to assay, HER18 cells were harvested using trypsin-EDTA, washed with phosphate-buffered saline (PBS) containing 5% serum and viable cells were counted. For culture of primary target cells, peripheral blood from CLL patients was subject to density centrifugation to obtain peripheral blood mononuclear cells (PBMC). Cell preparations were washed and then subject to positive selection of CD5 ⁺ CD19 ⁺ CLL cells using magnetic beads (MACS, Miltenyi Biotech). Cell preparations were used fresh after selection. NK cells from CLL patients and healthy volunteers were enriched from peripheral blood collected in sodium citrate anti-coagulant tubes and then subject to density centrifugation. Removal of non NK cells was performed using negative selection by MACS separation. Freshly isolated NK cells were washed three times, enumerated, and then used immediately for ADCC assays.

[00687] Cytokine secretion was determined as follows. Rituximab and trastuzumab-dependent NK-cell mediated degranulation and cytokine release were assessed using lymphoma and HER2+ breast cancer cell lines (DHL-4 and HER18, respectively). Target cells were cultured in flat-bottom plates containing 10 μg/mL of rituximab (DHL-4) or trastuzumab (HER18) and test articles (01 or 1 μM ibrutinib 1 μM Formula (2) or DMSO vehicle control) NK cells from healthy donors were enriched as described above and then added to the target cells and incubated for 4 hours at 37 ºC. Triplicate cultures were performed on NK cells from donors. After incubation, supernatants were harvested, centrifuged briefly, and then analyzed for interferon-γ using an enzyme-linked immunosorbent assay (ELISA) (R&D Systems, Minneapolis, MN, USA).

[00688] Lytic granule release was determined as follows. NK cells from healthy donors were enriched and cultured in the presence of target cells, monoclonal antibodies and test articles as described above. After 4 hours, the cultures were harvested and cells were pelleted, washed, and then stained for flow cytometry evaluation. Degranulation was evaluated via by flow cytometery by externalization of CD107a, a protein normally present on the inner leaflet of lytic granules, and gating on NK cells (CD3-CD16 ⁺ lymphocytes). The percentage of CD107a positive NK cells was quantified by comparison with a negative control (isotype control, unstained cells/FMO). Control cultures (NK cells cultured without target cells, or NK, target cell co- cultures in the absence of appropriate monoclonal antibody) were also evaluated; all experiments were performed in triplicate.

[00689] ADCC assays were performed as follows. Briefly, target cells (Raji or primary CLL) were labeled by incubation at 37 ºC with 100 μCi ⁵¹ Cr for 4 hours prior to co-culture with NK cells. Cells were washed, enumerated, and then added in triplicate to prepared 96-well plates containing treated NK cells at an effector:target (E:T) ratio of 25:1. Rituximab (Genentech) was added to ADCC wells at concentrations of 0.1, 1.0 or 10 μg/mL and the assays were briefly mixed and then centrifuged to collect cells at the bottom of the wells. The effect of NK cell natural cytotoxicity was assessed in wells containing no rituximab. Cultures were incubated at 37 ºC for 4 hours, and then centrifuged. Supernatants were harvested and ⁵¹ Cr release was measured by liquid scintillation counting. All experiments were performed in triplicate.

[00690] Ibrutinib inhibited rituximab-induced NK cell cytokine secretion in a dose-dependent manner (0.1 and 1 μM) (FIG.33: 48% p = 0.018; 72% p = 0.002, respectively). At 1 μM, Formula (2) did not significantly inhibit cytokine secretion (FIG.33: 3.5%). Similarly, Formula (2) had no inhibitory effect on rituximab-stimulated NK cell degranulation (< 2%) while ibrutinib reduced degranulation by ~50% (p = 0.24, FIG.34). Formula (2) had no inhibitory effect while ibrutinib prevented trastuzumab-stimulated NK cell cytokine release and degranulation by ~92% and ~84% at 1 μM, respectively (FIG.33 and FIG.34: ***p = 0.004, **p = 0.002).

[00691] In Raji cell samples, ex vivo NK cell activity against autologous tumor cells was not inhibited by addition of Formula (2) at 1 μM, and increased cell lysis was observed with increasing concentrations of rituximab at a constant E:T ratio (FIG.35). In primary CLL samples, ex vivo NK cell activity against autologous tumor cells was not inhibited by addition of Formula (2) at 1 μM, and increased cell lysis was observed with increasing concentrations of rituximab at a constant E:T ratio (FIG.36). In contrast, addition of 1 μM ibrutinib completely inhibited ADCC, with less than 10% cell lysis at any rituximab concentration and no increase in cell lysis in the presence of rituximab, compared with cultures without rituximab. A graph highlighting the significant differences between Formula (2) and ibrutinib is shown in FIG.37 (where“Ab” refers to rituximab). The difference between Formula (2) and ibrutinib was highly significant in this assay (p = 0.001).

[00692] In ADCC assays using healthy donor NK cells, antibody-dependent lysis of rituximab- coated Raji cells was not inhibited by addition of 1 μM of Formula (2) (FIG.37). In these experiments, addition of rituximab stimulated a 5- to 8-fold increase in cell lysis at 0.1 and 1 μg/mL, compared with low (< 20%) natural cytotoxicity in the absence of rituximab. As previously reported, addition of 1 μM ibrutinib strongly inhibited the antibody-dependent lysis of target cells, with less than 20% cell lysis at all rituximab concentrations and no increase in ADCC with at higher rituximab concentrations.

[00693] Ibrutinib is clinically effective as monotherapy and in combination with rituximab, despite inhibition of ADCC in vitro and in vivo murine models due to ibrutinib’s secondary irreversible binding to ITK. Preclinically, the efficacy of therapeutics which do not inhibit NK cell function, including Formula (2), is superior to ibrutinib. These results provide support for the unexpected property of Formula (2) as a synergistic and superior active pharmaceutical ingredient than ibrutinib to use in combinations with antibodies that have ADCC as a mechanism of action. The improved performance of Formula (2) in combination with anti-CD20 antibody therapies is expected to extend to its use in combination with CD19 and PD-1/PD-L1 inhibitors in both hematological malignancies and solid tumors, as these combinations would also benefit from reduced inhibition of NK cell function. The results also indicate that Formula (2) can be combined synergistically with anti-CD19 NK cell therapies, such as NK cells expressing CD19- targeted chimeric antigen receptors, while Formula (10) cannot be similarly combined without loss of function for NK cell therapies.

Example 8– Effects of BTK Inhibition on Antibody-Dependent NK Cell Mediated Cytotoxicity Using Obinutuzumab

[00694] It has been shown above that ibrutinib undesirably antagonizes rituximab ADCC effects mediated by NK cells, and that Formula (2) does not antagonize rituximab ADCC effects and instead allows for a synergistic combination. As noted previously, this may be due to ibrutinib’s secondary irreversible binding to ITK, which is required for FcR-stimulated NK cell function including calcium mobilization, granule release, and overall ADCC. H. E. Kohrt, et al., Blood 2014, 123, 1957-60. The potential for ibrutinib antagonization of obinutuzumab (GA-101) ADCC as mediated by NK cells was also explored and compared to the effects of Formula (2).

[00695] The NK cell degranulation/ADCC assay was performed using a whole blood assay with CLL targets added to normal donor whole blood, in the presence or absence of different doses of Formula (2) and ibrutinib, followed by opsonization with the anti-CD20 antibody obinutuzumab. Ibrutinib was used as a control, and two blinded samples of BTK inhibitors, Formula (2) and a second sample of ibrutinib, were provided to the investigators. Degranulation in whole blood was performed as follows. CLL targets (MEC-1 cells) were expanded in RPMI 1640 medium (Life Technologies, Inc.) with 10% fetal bovine serum (FBS). Exponentially growing cells were used. On the day of the experiment, 8 mL of blood was drawn from a normal volunteer into a test tube containing desirudin to obtain a final concentration of 50 μg/mL. A white blood cell (WBC) count of whole blood was performed. MEC-1 cells were re-suspended at the

concentration of WBC in whole blood (e.g., if 6 ×10 ⁶ WBC/mL was measured, MEC-1 cells were re-suspended at 6 ×10 ⁶ cells/mL, to allow for a final WBC:MEC-1 cell ratio of 1:1). The ibrutinib control and two blinded BTK inhibitors were diluted in X-VIVO 15 serum-free hematopoietic cell medium (Lonza Group, Ltd.) to concentrations of 200 μM, 20 μM and 2 μM. 170 μL aliquots of unmanipulated whole blood were incubated with 10 μL BTK inhibitors or X- VIVO 15 medium for one hour into a plate. Cetuximab and obinutuzumab (GA-101) were diluted in X-VIVO 15 medium to a concentration of 20 μg/mL. Equal volumes of MEC-1 cells and antibodies were incubated for 5 minutes. After incubation, 20 μL of MEC-1 cells and antibodies was added to whole blood and the BTK inhibitors/X-VIVO 15 medium (for a final volume of 200 μL). The samples were placed in a 5% CO ₂ incubator for 4 hours at 37 °C. The experimental conditions thus achieved a WBC:MEC-1 cell ratio of 1:1, with final concentrations of the BTK inhibitors in the assay of 10 μM, 1 μM and 0.1 μM and final concentrations of the antibodies of 1 μg/mL.

[00696] After 4 hours, the samples were mixed gently and 50 μL aliquots were removed from each well and placed in fluorescence-activated cell sorting (FACS) test tubes. A 20 μL aliquot of anti-CD56-APC antibody and anti-CD107a-PE antibody was added. The samples were incubated for 20 minutes at room temperature in the dark. An aliquot of 2 mL of FACS lysing solution (BD Biosceinces) was added. The samples were again incubated for 5 minutes, and then centrifuged at 2000 rpm for 5 minutes. Supernatant was discarded and the cell pellet was resuspended in 500 μL of PBS. The samples were analyzed on the flow cytometer for CD107a ⁺ NK cells (CD56 ⁺ ).

[00697] The NK cell degranulation results are summarized in FIG.38 for n = 3 experiments, which shows the effects on whole blood after pretreatment for 1 hour with the BTK inhibitors at the concentrations shown and subsequent stimulation with MEC-1 opsonised with obinutuzumab or cetuximab at 1 µg/mL for 4 hours. A strong reduction in the percentage of CD56 ⁺ /CD107a ⁺ NK cells is observed using ibrutinib (both as a control and blinded BTK inhibitor), which indicates that ibrutinib undesirably antagonizes NK cells. In contrast, Formula (2) shows little antagonism towards NK cells, and had a minimal effect on obinutuzumab-stimulated NK cell degranulation while ibrutinib reduced obinutuzumab-stimulated NK degranulation by greater than 40%. These results support the synergistic combination of obinutuzumab and Formula (2) in treatment of human B cell malignancies.

Example 9– Effects of BTK Inhibition on Generalized NK Cell Mediated Cytotoxicity

[00698] An assay was performed to assess the effects of BTK inhibition using Formula (2) on generalized NK killing (non-ADCC killing). The targets (K562 cells) do not express major histocompatibility complex (MHC) class I, so they do not inactivate NK cells. Target cells were grown to mid-log phase, and 5×10 ⁵ cells were labeled in 100 µL of assay medium (IMDM with 10% FCS and penicillin/streptomycin) with 100 µCi of ⁵¹ Cr for 1 hour at 37 °C. Cells were washed twice and resuspended in assay medium. A total of 5000 target cells/well was used in the assay. Effector cells were resuspended in assay medium, distributed on a V-bottom 96-well plate, and mixed with labeled target cells at 40:1 E:T ratios. Maximum release was determined by incubating target cells in 1% Triton X-100. For spontaneous release, targets were incubated without effectors in assay medium alone. After a 1 minute centrifugation at 1000 rpm, plates were incubated for 4 and 16 hours at 37 °C. Supernatant was harvested and ⁵¹ Cr release was measured in a gamma counter. Percentage of specific release was calculated as (experimental release-spontaneous release)/(maximum release-spontaneous release) × 100. The results are shown in FIG.39.

Example 10– Effects of BTK Inhibition on T Cells

[00699] An assay was performed to assess the effects of BTK inhibition using Formula (2) on T cells. Enriched CD4 ⁺ T cells are plated on 24-well culture dishes that have been precoated 2 hr with 250 μL anti-TCRβ (0.5 μg/mL) plus anti-CD28 (5 μg/mL) at 37 °C in PBS. The cells are then supplemented with media containing BTK inhibitors along with the skewing cytokines as indicated in the following. The Th17 and Treg cultures are grown for 4 days before analysis. The cells are maintained for an additional 3 days with skewing cytokines (Th17; 20 ng/mL IL-6, 0.5 ng/mL TGF-β, 5 μg/mL IL-4, 5 μg/mL IFN-γ and Treg; 0.5 ng/mL TGF-β, 5 μg/mL IL-4, 5 μg/mL IFN-γ) and are supplemented with IL2 as a growth factor.

[00700] The results are shown in FIG.40 and FIG.41, and further illustrate the surprising properties of Formula (2) in comparison to Formula (10) (ibrutinib). Because of the lack of activity of Formula (2) on ITK and Txk, no adverse effect on Th17 and Treg development was observed. Since ibrutinib inhibits both ITK and Txk, a profound inhibition of Th17 cells and an increase in Treg development is observed, which is comparable to the murine ITK/Txk double knock-out cells which were used as a control.

[00701] The effects of ibrutinib in comparison to Formula (2) on CD8 ⁺ T cell viability were also assessed. Total T cells were plated on anti-TCR and anti-CD28 coated wells in the presence of both BTK inhibitors. Neutral culture conditions were used that will not polarize T cells to a helper lineage. The cells are grown for 4 days and are then stained with anti-CD4, anti-CD8 and LIVE/DEAD reagent to determine if the drugs have selective effects on either the CD4 ⁺ or CD8 ⁺ cells. Statistical significance was calculated using the Mann Whitney T-test. The results, shown in FIG.42, indicate that higher concentrations of Formula (10) (ibrutinib) have a strong, negative effect on CD8 ⁺ T cell viability that is not observed with Formula (2) at any concentration. [00702] Without being bound by any theory, CD8 ⁺ T cells have two primary effector functions: (1) produce large amounts of IFN- ^ (which activates macrophages), and (2) cytolytic activity. A cytotoxic T cell (CTL) assay was performed to compare the BTK inhibitors of Formula (2) and Formula (10) (ibrutinib). Effectors were prepared by generating CTL by culturing MHC mismatched splenocytes for 4 days with (500 nM) and without Formula (2) or Formula (10) (ibrutinib). The targets were B lymphoblasts from lipopolysaccharide (LPS) treated cultures. The assay was performed by incubating different ratios of effectors:targets for 4 hours. In FIG. 43, the results show that Formula (10) (ibrutinib) affects CD8 ⁺ T cell function as measured by % cytotoxicity. Formula (2), in contrast, has no effect on CD8 ⁺ T cell function as measured by % cytotoxicity relative to vehicle. The effect on CD8 ⁺ T cell function can also be observed by measurement of IFN- ^ levels, as shown in FIG.44, where Formula (10) (ibrutinib) again results in a significant loss of function relative to Formula (2) and vehicle.

[00703] As described herein, the tumor microenvironment provides stromal support for tumor growth and conceals the tumor from immune surveillance. Tumor-associated stroma comprises a mix of fibroblasts, Tregs, MDSCs, and TAMs that promote tumor growth and restrain immune- mediated tumor cell killing. The targeting of immune infiltrates may impair stromal support and enhance immune-mediated killing of cancer cells. Specifically, myeloid cells significantly infiltrate the tumor microenvironment and suppress anti-tumor immunity, promote tumor growth and tumor metastasis. Two subsets of myeloid cells with significant immunosuppressive activity are MDSCs and TAMs. During tumor progression, MDSCs and TAMs facilitate tumor growth by down regulating the host immune response through suppression of CD8 ⁺ T cell activation and function. BTK is expressed among cells of hematopoietic origin including B cells, myeloid cells, mast cells and platelets, but not T cells, where it regulates multiple cellular processes. BTK-dependent activation of myeloid cells and lymphocytes sustain the complex

microenvironment needed for lymphoid and solid tumor maintenance. BTK inhibition shifts the microenvironment from suppressive to stimulatory by inhibiting the infiltration and function and of innate immune subsets (MDSCs and TAMs), thus allowing the activation of cytotoxic CD8 ⁺ T cells. The unique selectivity of the BTK inhibitor of Formula (2), as compared to, e.g., the BTK inhibitor of Formula (10) (ibrutinib), prevents loss of function for CD8 ⁺ T cells as shown in this example. Further evidence of this effect is provided by the results shown in FIG.9 and FIG.31, with the T cell promoting effects of Formula (2) further highlighted e g in FIG 25 As a result T cell therapies including CAR-T (CD19-targeted chimeric antigen receptor expressing CD8 ⁺ T cells) can be used in combination with Formula (2) and other selective BTK inhibitors described herein in a synergistic manner. Formula (10) cannot be coadministered without loss of function for CD8 ⁺ CD19-targeted chimeric antigen receptor expressing T cells.

[00704] Recent clinical trials have shown that CAR-T cells are a promising therapy for hematological and solid tumor malignancies. CARs include fusion proteins combining the antigen-recognition fragment of a monoclonal antibody with T cell activation domains from the T-cell receptor complex, such as the ζ chain, and costimulatory endodomains, from CD28, 4- 1BB, or OX40. In clinical trials, up to 90% complete response rates have been seen after CD19- CAR-T administration, even in chemotherapy-refractory acute lymphocytic leukemia. Results in other B-cell cancers, such as chronic lymphocytic leukemia (CLL) and diffuse large B-cell lymphoma (DLBCL), however, have been less striking. One explanation for the different response rates among tumor types is that CAR-T functionality may be inhibited by an immunosuppressive tumor microenvironment. As mentioned above, TAMs, macrophages, and MDSCs inhibit T cell responses and promote recruitment of regulatory T cells. Thus, a BTK inhibitor that promotes T cell function (such as Formula (2)) may have diverse and synergistic benefits in combination with adoptive T cell therapy and CAR-T cells in clinical practice through reducing the inhibitory tumor microenvironment by inhibiting the infiltration and function and of innate immune subsets in the tumor microenvironment.

Example 11– Clinical Study of a BTK Inhibitor in Leukemia/Lymphoma and Effects on Bone Marrow and Lymphoid Microenvironments

[00705] Clinical studies have shown that targeting the BCR signaling pathway by inhibiting BTK produces significant clinical benefit in patients with non-Hodgkin’s lymphoma (NHL). The second generation BTK inhibitor, Formula (2), achieves significant oral bioavailability and potency, and has favorable preclinical characteristics, as described above. The purpose of this study is to evaluate the safety and efficacy of the second generation BTK inhibitor of Formula (2) in treating subjects with chronic lymphocytic leukemia (CLL) and small lymphocytic lymphoma (SLL).

[00706] The design and conduct of this study is supported by an understanding of the history and current therapies for subjects with lymphoid cancers; knowledge of the activity and safety of a first-generation BTK inhibitor ibrutinib in subjects with hematologic cancers; and the available nonclinical information regarding Formula (2). The collective data support the following conclusions. BTK expression plays an important role in the biology of lymphoid neoplasms, which represent serious and life-threatening disorders with continuing unmet medical need. Clinical evaluation of Formula (2) as a potential treatment for these disorders has sound scientific rationale based on observations that the compound selectively abrogates BTK activity and shows activity in nonclinical models of lymphoid cancers. These data are supported by clinical documentation that ibrutinib, a first-generation BTK inhibitor, is clinically active in these diseases. Ibrutinib clinical data and Formula (2) nonclinical safety pharmacology and toxicology studies support the safety of testing Formula (2) in subjects with B cell malignancies.

[00707] The primary objectives of the clinical study are as follows: (1) establish the safety and the MTD of orally administered Formula (2) in subjects with CLL/SLL; (2) determine pharmacokinetics (PK) of orally administered Formula (2) and identification of its major metabolite(s); and (3) measure pharmacodynamic (PD) parameters including drug occupancy of BTK, the target enzyme, and effect on biologic markers of B cell function.

[00708] The secondary objective of the clinical study is to evaluate tumor responses in patients treated with Formula (2).

[00709] This study is a multicenter, open-label, nonrandomized, sequential group, dose escalation study. The following dose cohorts will be evaluated:

Cohort 1: 100 mg/day for 28 days (= 1 cycle)

Cohort 2: 175 mg/day for 28 days (= 1 cycle)

Cohort 3: 250 mg/day for 28 days (= 1 cycle)

Cohort 4: 350 mg/day for 28 days (= 1 cycle)

Cohort 5: 450 mg/day for 28 days (= 1 cycle)

Cohort 6: To be determined amount in mg/day for 28 days (= 1 cycle)

[00710] Each cohort will be enrolled sequentially with 6 subjects per cohort. If≤ 1 dose- limiting toxicity (DLT) is observed in the cohort during Cycle 1, escalation to the next cohort will proceed. Subjects may be enrolled in the next cohort if 4 of the 6 subjects enrolled in the cohort completed Cycle 1 without experiencing a DLT, while the remaining 2 subjects are completing evaluation. If≥ 2 DLTs are observed during Cycle 1, dosing at that dose and higher will be suspended and the MTD will be established as the previous cohort. The MTD is defined as the largest daily dose for which fewer than 33% of the subjects experience a DLT during Cycle 1. Dose escalation will end when either the MTD is achieved or at 3 dose levels above full BTK occupancy, whichever occurs first. Full BTK occupancy is defined as Formula (2) active- site occupancy of > 80% (average of all subjects in cohort) at 24 hours postdose. Should escalation to Cohort 6 be necessary, the dose will be determined based on the aggregate data from Cohorts 1 to 5, which includes safety, efficacy, and PK/PD results. The dose for Cohort 6 will not exceed 900 mg/day.

[00711] Treatment with Formula (2) may be continued for > 28 days until disease progression or an unacceptable drug-related toxicity occurs. Subjects with disease progression will be removed from the study. All subjects who discontinue study drug will have a safety follow-up visit 30 (±7) days after the last dose of study drug unless they have started another cancer therapy within that timeframe. Radiologic tumor assessment will be done at screening and at the end of Cycle 2, Cycle 4, and Cycle 12 and at investigator discretion. Confirmation of complete response (CR) will require bone marrow analysis and radiologic tumor assessment. For subjects who remain on study for > 11 months, a mandatory bone marrow aspirate and biopsy is required in Cycle 12 concurrent with the radiologic tumor assessment.

[00712] All subjects will have standard hematology, chemistry, and urinalysis safety panels done at screening. This study also includes pancreatic function assessment (serum amylase and serum lipase) due to the pancreatic findings in the 28-day GLP rat toxicity study. Once dosing commences, all subjects will be evaluated for safety once weekly for the first 4 weeks, every other week for Cycle 2, and monthly thereafter. Blood samples will be collected during the first week of treatment for PK/PD assessments. ECGs will be done at screening, and on Day 1-2, 8, 15, 22, 28 of Cycle 1, Day 15 and 28 of Cycle 2, and monthly thereafter through Cycle 6. ECGs are done in triplicate for screening only. Thereafter, single ECG tests are done unless a repeat ECG testing is required.

[00713] Dose-limiting toxicity is defined as any of the following events (if not related to disease progression): (1) any Grade≥ 3 non-hematologic toxicity (except alopecia) persisting despite receipt of a single course of standard outpatient symptomatic therapy (e.g., Grade 3 diarrhea that responds to a single, therapeutic dose of Imodium® would not be considered a DLT); (2) grade≥ 3 prolongation of the corrected QT interval (QTc), as determined by a central ECG laboratory overread; (3) grade 4 neutropenia (absolute neutrophil count [ANC] < 500/μL) lasting > 7 days after discontinuation of therapy without growth factors or lasting > 5 days after discontinuation of therapy while on growth factors (i.e., Grade 4 neutropenia not lasting as long as specified will not be considered a DLT), (4) grade 4 thrombocytopenia (platelet count < 20,000/μL) lasting > 7 days after discontinuation of therapy or requiring transfusion (i.e., Grade 4 thrombocytopenia not lasting as long as specified will not be considered a DLT), and (5) dosing delay due to toxicity for > 7 consecutive days.

[00714] The efficacy parameters for the study include overall response rate, duration of response, and progression-free survival (PFS). The safety parameters for the study include DLTs and MTD, frequency, severity, and attribution of adverse events (AEs) based on the Common Terminology Criteria for Adverse Events (CTCAE v4.03) for non-hematologic AEs. Hallek, et al., Blood 2008, 111, 5446-5456.

[00715] The schedule of assessments is as follows, with all days stated in the following meaning the given day or +/- 2 days from the given day. A physical examination, including vital signs and weight, are performed at screening, during cycle 1 at 1, 8, 15, 22, and 28 days, during cycle 2 at 15 and 28 days, during cycles 3 to 24 at 28 days, and at follow up (after the last dose). The screening physical examination includes, at a minimum, the general appearance of the subject, height (screening only) and weight, and examination of the skin, eyes, ears, nose, throat, lungs, heart, abdomen, extremities, musculoskeletal system, lymphatic system, and nervous system. Symptom-directed physical exams are done thereafter. Vital signs (blood pressure, pulse, respiratory rate, and temperature) are assessed after the subject has rested in the sitting position. Eastern Cooperative Oncology Group (ECOG) status is assessed at screening, during cycle 1 at 1, 8, 15, 22, and 28 days, during cycle 2 at 15 and 28 days, during cycles 3 to 24 at 28 days, and at follow up, using the published ECOG performance status indications described in Oken, et al., Am. J. Clin. Oncol.1982, 5, 649-655. ECG testing is performed at screening, during cycle 1 at 1, 2, 8, 15, 22, and 28 days, during cycle 2 at 15 and 28 days, during cycles 3 to 24 at 28 days, and at follow up. The 12-lead ECG test will be done in triplicate (≥ 1 minute apart) at screening. The calculated QTc average of the 3 ECGs must be <480 ms for eligibility. On cycle 1, day 1 and cycle 1 day 8 single ECGs are done predose and at 1 2 4 and 6 hours postdose The single ECG on Cycle 1 Day 2 is done predose. On cycle 1, day 15, day 22, and day 28, a single ECG is done 2 hours post-dose. Starting with cycle 2, a single ECG is done per visit. Subjects should be in supine position and resting for at least 10 minutes before study-related ECGs. Two consecutive machine-read QTc > 500 ms or > 60 ms above baseline require central ECG review. Hematology, including complete blood count with differential and platelet and reticulocyte counts, is assesed at screening, during cycle 1 at 1, 8, 15, 22, and 28 days, during cycle 2 at 15 and 28 days, during cycles 3 to 24 at 28 days, and at follow up. Serum chemistry is assesed at screening, during cycle 1 at 1, 8, 15, 22, and 28 days, during cycle 2 at 15 and 28 days, during cycles 3 to 24 at 28 days, and at follow up. Serum chemistry includes albumin, alkaline phosphatase, ALT, AST, bicarbonate, blood urea nitrogen (BUN), calcium, chloride, creatinine, glucose, lactate dehydrogenase (LDH), magnesium, phosphate, potassium, sodium, total bilirubin, total protein, and uric acid. Cell counts and serum immunoglobulin are performed at screening, at cycle 2, day 28, and at every 6 months thereafter until last dose and include T/B/NK/monocyte cell counts (CD3, CD4, CD8, CD14, CD19, CD19, CD16/56, and others as needed) and serum immunoglobulin (IgG, IgM, IgA, and total immunoglobulin). Bone marrow aspirates are performed at cycle 12. Pharmacodynamics samples are drawn during cycle 1 at 1, 2, and 8 days, and at follow up. On days 1 and 8, pharmacodynamic samples are drawn pre-dose and 4 hours (±10 minutes) post-dose, and on day 2, pharmacodynamic samples are drawn pre- dose. Pharmacokinetics samples are drawn during cycle 1 at 1, 2, 8, 15, 22, and 28 days.

Pharmacokinetic samples for Cycle 1 Day 1 are drawn pre-dose and at 0.5, 1, 2, 4, 6 and 24 hours (before dose on Day 2) post-dose. Samples for Cycle 1 Day 8 are drawn pre-dose and at 0.5, 1, 2, 4, and 6 hours post-dose. On Cycle 1 Day 15, 22, and 28, a PK sample is drawn pre- dose and the second PK sample must be drawn before (up to 10 minutes before) the ECG acquisition, which is 2 hours postdose. Pretreatment radiologic tumor assessments are performed within 30 days before the first dose. A computed tomography (CT) scan (with contrast unless contraindicated) is required of the chest, abdomen, and pelvis. In addition, a positron emission tomography (PET) or PET/CT must done for subjects with SLL. Radiologic tumor assessments are mandatory at the end of Cycle 2 (-7 days), Cycle 4 (- 7days), and Cycle 12 (-7 days).

Otherwise, radiologic tumor assessments are done at investigator discretion. A CT (with contrast unless contraindicated) scan of the chest, abdomen, and pelvis is required for subjects with CLL. In addition, a PET/CT is required in subjects with SLL. Bone marrow and radiologic assessments are both required for confirmation of a complete response (CR). Clinical assessments of tumor response should be done at the end of Cycle 6 and every 3 months thereafter. Molecular markers are measured at screening, and include interphase cytogenetics, stimulated karyotype, IgHV mutational status, Zap-70 methylation, and beta-2 microglobulin levels. Urinalysis is performed at screening, and includes pH, ketones, specific gravity, bilirubin, protein, blood, and glucose. Other assessments, including informed consent, eligibility, medical history, and pregnancy test are done at the time of screening.

[00716] The investigator rates the subject’s response to treatment based on recent guidelines for CLL, as given in Hallek, et al., Blood 2008, 111, 5446-56, and for SLL, as given in Cheson, et al., J. Clin. Oncol.2007, 25, 579-586. The response assessment criteria for CLL are summarized in Table 12.

TABLE 12. Response Assessment Criteria for CLL. Abbreviations: ANC = absolute neutrophil count; CR = complete remission; CRi = CR with incomplete blood count recovery; PR = partial remission.

a. ompute tomograpy scan o a omen, pevs, an cests requre or t s evauaton b. Without need for exogenous growth factors

c. In the sum products of≤ 6 lymph nodes or in the largest diameter of the enlarged lymph node(s) detected before therapy and no increase in any lymph node or new enlarged lymph nodes

[00717] The response assessment criteria for SLL are summarized in Table 13.

TABLE 13. Response Assessment Criteria for SLL. Abbreviations: CR = complete remission, CT = computed tomography, FDG = [ ¹⁸ F]fluorodeoxyglucose, PET = positron-emission tomography, PR = partial remission, SD = stable disease, SPD = sum of the product of the diameters.

[00718] The PK parameters of the study are as follows. The plasma PK of Formula (2) and a metabolite is characterized using noncompartmental analysis. The following PK parameters are calculated, whenever possible, from plasma concentrations of Formula (2): AUC _(0-t) : Area under the plasma concentration-time curve calculated using linear trapezoidal summation from time 0 to time t, where t is the time of the last measurable concentration (Ct),

AUC _(0-24) : Area under the plasma concentration-time curve from 0 to 24 hours, calculated using linear trapezoidal summation,

AUC _(0-∞) : Area under the plasma concentration-time curve from 0 to infinity, calculated using the formula: AUC _(0-∞) = AUC _(0-t) + Ct / λz, where λz is the apparent terminal elimination rate constant,

C _max : Maximum observed plasma concentration,

T _max : Time of the maximum plasma concentration (obtained without interpolation), t _½ : Terminal elimination half-life (whenever possible),

λ _z : Terminal elimination rate constant (whenever possible),

Cl/F: Oral clearance.

[00719] The PD parameters of the study are as follows. The occupancy of BTK by Formula (2) are measured in peripheral blood mononuclear cells (PBMCs) with the aid of a biotin-tagged Formula (2) analogue probe. The effect of Formula (2) on biologic markers of B cell function will also be evaluated.

[00720] The statistical analysis used in the study is as follows. No formal statistical tests of hypotheses are performed. Descriptive statistics (including means, standard deviations, and medians for continuous variables and proportions for discrete variables) are used to summarize data as appropriate.

[00721] The following definitions are used for the safety and efficacy analysis sets: Safety analysis set: All enrolled subjects who receive≥ 1 dose of study drug; Per-protocol (PP) analysis set: All enrolled subjects who receive≥ 1 dose of study drug and with≥ 1 tumor response assessment after treatment. The safety analysis set will be used for evaluating the safety parameters in this study. The PP analysis sets will be analyzed for efficacy parameters in this study.

[00722] No imputation of values for missing data is performed except for missing or partial start and end dates for adverse events and concomitant medication will be imputed according to prespecified, conservative imputation rules. Subjects lost to follow-up (or drop out) will be included in statistical analyses to the point of their last evaluation.

[00723] The safety endpoint analysis was performed as follows. Safety summaries will include summaries in the form of tables and listings. The frequency (number and percentage) of treatment emergent adverse events will be reported in each treatment group by Medical

Dictionary for Regulatory Activities (MedDRA) System Organ Class and Preferred Term.

Summaries will also be presented by the severity of the adverse event and by relationship to study drug. Laboratory shift tables containing counts and percentages will be prepared by treatment assignment, laboratory parameter, and time. Summary tables will be prepared for each laboratory parameter. Figures of changes in laboratory parameters over time will be generated. Vital signs, ECGs, and physical exams will be tabulated and summarized.

[00724] Additional analyses include summaries of subject demographics, baseline

characteristics, compliance, and concurrent treatments. Concomitant medications will be coded according to the World Health Organization (WHO) Drug Dictionary and tabulated.

[00725] The analysis of efficacy parameters was performed as follows. The point estimate of the overall response rate will be calculated for the PP analysis set. The corresponding 95% confidence interval also will be derived. The duration of overall response is measured from the time measurement criteria are met for CR or PR (whichever is first recorded) until the first date that recurrent or progressive disease is objectively documented (taking as reference for progressive disease the smallest measurements recorded since the treatment started). Kaplan- Meier methodology will be used to estimate event-free curves and corresponding quantiles (including the median). Progression-free survival is measured from the time of first study drug administration until the first date that recurrent or progressive disease is objectively documented (taking as reference for progressive disease the smallest measurements recorded since the treatment started). Kaplan-Meier methodology will be used to estimate the event-free curves and corresponding quantiles (including the median).

[00726] The study scheme is a sequential cohort escalation. Each cohort consists of six subjects. The sample size of the study is 24 to 36 subjects, depending on dose escalation into subsequent cohorts. Cohort 1 (N = 6) consists of Formula (2), 100 mg QD for 28 days. Cohort 2 (N = 6) consists of Formula (2), 175 mg QD for 28 days. Cohort 3 (N = 6) consists of Formula (2), 250 mg QD for 28 days. Cohort 4 (N = 6) consists of Formula (2), 350 mg QD for 28 days. Cohort 5 (N = 6) consists of Formula (2), 450 mg QD for 28 days. Cohort 6 (N = 6) consists of Formula (2), at a dose to be determined QD for 28 days. The dose level for Cohort 6 will be determined based on the safety and efficacy of Cohorts 1 to 5, and will not exceed 900 mg/day. Escalation will end with either the MTD cohort or three levels above full BTK occupancy, whichever is observed first. An additional arm of the study will explore 100 mg BID dosing. Treatment with oral Formula (2) may be continued for greater than 28 days until disease progression or an unacceptable drug-related toxicity occurs.

[00727] The inclusion criteria for the study are as follows: (1) men and women≥ 18 years of age with a confirmed diagnosis of CLL/SLL, which has relapsed after, or been refractory to,≥ 2 previous treatments for CLL/SLL; however, subjects with 17p deletion are eligible if they have relapsed after, or been refractory to, 1 prior treatment for CLL/SLL; (2) body weight≥ 60 kg, (3) ECOG performance status of≤ 2; (4) agreement to use contraception during the study and for 30 days after the last dose of study drug if sexually active and able to bear children; (5) willing and able to participate in all required evaluations and procedures in this study protocol including swallowing capsules without difficulty; or (6) ability to understand the purpose and risks of the study and provide signed and dated informed consent and authorization to use protected health information (in accordance with national and local subject privacy regulations).

[00728] The dosage form and strength of Formula (2) used in the clinical study is a hard gelatin capsules prepared using standard pharmaceutical grade excipients (microcrystalline cellulose) and containing 25 mg of Formula (2) each. The color of the capsules is Swedish orange. The route of administration is oral (per os, or PO). The dose regimen is once daily or twice daily, as defined by the cohort, on an empty stomach (defined as no food 2 hours before and 30 minutes after dosing).

[00729] The baseline characteristics for the patients enrolled in the clinical study are given in Table 14.

TABLE 14. Relapsed/refractory CLL baseline characteristics.

[00730] The results of the clinical study in relapsed/refractory CLL patients are summarized in Table 15.

TABLE 15. Activity of Formula (2) in relapsed/refractory CLL. (PR = partial response; PR+L = partial response with lymphocytosis; SD = stable disease; PD = progressive disease.)

[00731] FIG.45 shows the median % change in ALC and SPD from baseline in the clinical study of Formula (2), plotted in comparison to the results reported for ibrutinib in Figure 1A of Byrd, et al., N. Engl. J. Med.2013, 369, 32-42. The results show that Formula (2) leads to a more rapid patient response in CLL than corresponding treatment with ibrutinib. This effect is illustrated, for example, by the median % change in SPD, which achieved the same status in the present study at 7 months of treatment with Formula (2) as compared to 18 months for ibrutinib. The % change in SPD observed in the different cohorts (i.e. by dose and dosing regimen) is shown in FIG.46, and in all cases shows significant responses.

[00732] A Kaplan-Meier curve showing PFS from the clinical CLL study of Formula (2) is shown in FIG.47. A comparison of survival curves was performed using the Log-Rank (Mantle- Cox) test, with a p-value of 0.0206 indicating that the survival curves are different. The number of patients at risk is shown in FIG.48. Both FIG.47 and FIG.48 show the results for Formula (2) in comparison to the results reported for Formula (10) (ibrutinib) in Byrd, et al., N. Engl. J. Med.2013, 369, 32-42. An improvement in survival and a reduction in risk are observed in CLL patients treated with Formula (2) in comparison to patients treated with ibrutinib.

[00733] Based on the data and comparisons shown above, the CLL study showed that the efficacy of Formula (2) was surprisingly superior to that of Formula (10) (ibrutinib).

[00734] In the literature study of ibrutinib, increased disease progression was associated with patients with high-risk cytogenetic lesions (17p13.1 deletion or 11q22.3 deletion), as shown in Figure 3A in Byrd, et al., N. Engl. J. Med.2013, 369, 32-42, which shows ibrutinib PFS including PFS broken down by genetic abnormality. The 17p and 11q deletions are validated high-risk characteristics of CLL, and the 17p deletion is the highest risk. In FIG.49, the PFS is shown for Formula (2) in patients with the 17p deletion in comparison to the results obtained for ibrutinib in Byrd, et al., N. Engl. J. Med.2013, 369, 32-42. A p-value of 0.0696 was obtained. In FIG.50, the number of patients at risk with the 17p deletion is compared. To date, no 17p patients have progressed on Formula (2).

[00735] The adverse events observed in the clinical study in relapsed/refractory CLL are given in Table 16. No DLTs were observed. The MTD was not reached. No treatment-related serious adverse events (SAEs) were observed. No prophylactic antivirals or antibiotics were needed. TABLE 16. Treatment-related adverse events reported in the clinical study of Formula (2) in relapsed/refractory CLL. (Reported in≥ 5% of patients.)

[00736] The clinical study of Formula (2) thus showed other unexpectedly superior results compared to ibrutinib therapy. A lack of lymphocytosis was observed in the study.

Furthermore, only grade 1 AEs were observed, and these AEs were attributable to the high BTK selectivity of Formula (2).

[00737] BTK target occupancy was measured for relapsed/refractory CLL patients with the results shown in FIG.51. For 200 mg QD dosing of the BTK inhibitor of Formula (2), about 94% - 99% BTK occupancy was observed, with superior 24 hour coverage and less inter-patient variability also observed. For 420 mg and 840 mg QD of the BTK inhibitor ibrutinib, 80% - 90% BTK occupancy was observed, with more inter-patient variability and capped occupancy. These results indicate that the BTK inhibitor of Formula (2) achieves superior BTK occupancy in CLL patients than ibrutinib.

[00738] The effects of Formula (2) on cell subset percentages were also evaluated using flow cytometry analysis of peripheral blood, with the results shown in FIG.52, FIG.53, FIG.54, FIG. 55, FIG.56, and FIG.57. PBMC samples from CLL patient samples drawn prior to (predose) and after 28 days of dosing with Formula (2) were compared for potential changes in cell subsets. PBMCs were stained with monoclonal antibodies conjugated to fluorescent tags (flourochromes) to identify cell subsets via flow cytometry. Non-viable cells were excluded from the analysis using the dye 7-aminoactinomycin D (7-AAD). To produce the metric of percent change, the following steps were taken. First, each cell subset was defined by hierarchical flow cytometry gating. Then, the change in frequency (between day 1 and day 28) was calculated for each cell subset. MDSC subsets were measured as a % of all myeloid cells. T cell subsets were measured as a % of all CD3 ⁺ cells, and NK cells were measured as a % of all live CD45 ⁺ cells. In FIG.52 and FIG.53, the results show the % change in MDSC (monocytic) level over 28 days versus % ALC change at cycle 1 day 28 (C1D28) and at cycle 2 day 28 (C2D28). A cycle is 28 days. A trend is observed wherein patients with decreasing ALC % had increasing MDSC (monocytic) %. This may include patients who had quickly resolving lymphocytosis and those with no initial lymphocytosis. This provides evidence that treatment with Formula (2) mobilizes MDSCs and thus affects the CLL tumor microenvironment in marrow and lymph nodes, which is an unexpected indication of superior efficacy. In FIG.54 and FIG.55, the results show the % change in NK cell level over 28 days versus % ALC change, measured at C1D28 or C2D28, and similar trends are observed wherein patients with decreasing ALC % had increasing NK cell %. This may include patients who had quickly resolving lymphocytosis and those having no initial lymphocytosis. The effects in FIG.52 to FIG.55 are observed in multiple cohorts, at doses including 100 mg BID, 200 mg QD, and 400 mg QD. In FIG.56 and FIG.57, the effects on NK cells and MDSC cells are compared to a number of other markers versus % change in ALC at C1D28 and C2D28. These other markers include CD4+ T cells, CD8+ T cells, CD4+/CD8+ T cell ratio, NK-T cells, PD-1+ CD4+ T cells, and PD-1+ CD8+ T cells. The effects on NK cells and MDSC cells are observed to be much more pronounced than on any of these other markers.

[00739] These results suggest that after Formula (2) administration, the CLL microenvironment undergoes a change wherein NK cells and monocytic MDSC subsets increase in frequency in the peripheral blood in patients with falling ALC counts, an important clinical parameter in CLL. The NK cell increase may reflect an overall increase in cytolytic activity against B-CLL resulting in the ALC % to drop. The increase in MDSC % in the blood may be due to a movement of these cells out of the lymph nodes, spleen, and bone marrow, which are all possible sites of CLL proliferation. Fewer MDSCs at the CLL proliferation centers would likely result in a reduced immunosuppressive microenvironment leading to an increase in cell-mediated immunity against the tumor, decreased tumor proliferation, and eventually lower ALC% in the circulation.

[00740] Updated clinical results from the CLL study are shown in FIG.58 to FIG.63. FIG.58 shows an update of the data presented in FIG.45. FIG.59 shows an update of the data presented in FIG.51, and includes BID dosing results. Formula (2) 200 mg QD dosing resulted in 94% - 99% BTK occupancy, 24 hour coverage, and less inter-patient variability. Ibrutinib 420 mg and 840 mg QD dosing resulted in 80% - 90% BTK occupancy, more inter-patient variability, and capped occupancy Formula (2) 100 mg BID dosing resulted in 97% 99% BTK occupancy complete BTK coverage, and less inter-patient variability. The PFS for patients with 17p deletions and 11q deletions are illustrated in FIG.60, FIG.61, and FIG.62. Updated SPD results are illustrated in FIG.63, and again show significant results across all cohorts and dosing regimens.

[00741] Treatment of CLL patients with Formula (2) also resulted in increased apoptosis, as illustrated in FIG.64. Apoptotic B-CLL was defined by flow cytometry as having cleaved PARP ⁺ , Caspase 3 ⁺ , CD19 ⁺ , and CD5 ⁺ phenotypes. 82% of samples tested had a baseline change greater than 25%. Treatment of CLL patients also showed that Formula (2) decreased plasma chemokines associated with MDSC homing and retention. A significant decrease in CXCL12 and CCL2 levels has been observed in patients treated with Formula (2), as shown in FIG.65 and FIG.66, respectively.

[00742]

Example 12– Effects of BTK Inhibitors on Thrombosis

[00743] Clinical studies have shown that targeting the BCR signaling pathway by inhibiting BTK produces significant clinical benefit (Byrd, et al., N. Engl. J. Med.2013, 369(1), 32-42, Wang, et al., N. Engl. J. Med.2013, 369(6), 507-16). However, in these studies, bleeding has been reported in up to 50% of ibrutinib-treated patients. Most bleeding events were of grade 1-2 (spontaneous bruising or petechiae) but, in 5% of patients, they were of grade 3 or higher after trauma. These results are reflected in the prescribing information for ibrutinib, where bleeding events of any grade, including bruising and petechiae, were reported in about half of patients treated with ibrutinib (IMBRUVICA package insert and prescribing information, revised July 2014, U.S. Food and Drug Administration).

[00744] Constitutive or aberrant activation of the BCR signaling cascade has been implicated in the propagation and maintenance of a variety of B cell malignancies. Small molecule inhibitors of BTK, a protein early in this cascade and specifically expressed in B cells, have emerged as a new class of targeted agents. There are several BTK inhibitors, including Formula (17) (CC- 292), and Formula (10) (ibrutinib), in clinical development. Importantly, early stage clinical trials have found ibrutinib to be particularly active in chronic lymphocytic leukemia (CLL) and mantle cell lymphoma (MCL), suggesting that this class of inhibitors may play a significant role in various types of cancers (Aalipour and Advani, Br. J. Haematol.2013, 163, 436-43). However, their effects are not limited to leukemia or lymphomas as platelets also rely on the Tec kinases family members BTK and Tec for signal transduction in response to various

thrombogenic stimuli (Oda, et al., Blood 2000, 95(5), 1663-70; Atkinson, et al., Blood 2003, 102(10), 3592-99). In fact, both Tec and BTK play an important role in the regulation of phospholipase Cγ2 (PLCγ2) downstream of the collagen receptor glycoprotein VI (GPVI) in human platelets. In addition, BTK is activated and undergoes tyrosine phosphorylation upon challenge of the platelet thrombin receptor, which requires the engagement of αIIbβ3 integrin and PI3K activity (Laffargue, et al., FEBS Lett.1999, 443(1), 66-70). It has also been implicated in GPIbα-dependent thrombus stability at sites of vascular injury (Liu, et al., Blood 2006, 108(8), 2596-603). Thus, BTK and Tec are involved in several processes important in supporting the formation of a stable hemostatic plug, which is critical for preventing significant blood loss in response to vascular injury. Hence, the effects of the BTK inhibitors of Formula (2) and

Formula (10) (ibrutinib) were evaluated on human platelet-mediated thrombosis by utilizing the in vivo human thrombus formation in the VWF HA1 mice model described in Chen, et al., Nat. Biotechnol.2008, 26(1), 114-19.

[00745] Administration of anesthesia, insertion of venous and arterial catheters, fluorescent labeling and administration of human platelets (5 × 10 ⁸ /ml), and surgical preparation of the cremaster muscle in mice have been previously described (Chen et al., Nat Biotechnol.2008, 26(1), 114-19). Injury to the vessel wall of arterioles (~40–65 mm diameter) was performed using a pulsed nitrogen dye laser (440 nm, Photonic Instruments) applied through a 20× water- immersion Olympus objective (LUMPlanFl, 0.5 numerical aperture (NA)) of a Zeiss Axiotech vario microscope. Human platelet and wall interactions were visualized by fluorescence microscopy using a system equipped with a Yokogawa CSU-22 spinning disk confocal scanner, iXON EM camera, and 488 nm and 561 nm laser lines to detect BCECF-labeled and rhodamine- labeled platelets, respectively (Revolution XD, Andor Technology). The extent of thrombus formation was assessed for 2 minutes after injury and the area (μm ² ) of coverage determined (Image IQ, Andor Technology). For the Formula (2), Formula (17) (CC-292), and Formula (10) (ibrutinib) inhibition studies, the BTK inhibitors were added to purified human platelets for 30 minutes before administration.

[00746] The in vivo throbus effects of the BTK inhibitors, Formula (2), Formula (17) (CC-292), and Formula (10) (ibrutinib) were evaluated on human platelet mediated thrombosis by utilizing the in vivo human thrombus formation in the VWF HA1 mice model, which has been previously described (Chen, et al., Nat Biotechnol.2008, 26(1), 114-19). Purified human platelets were preincubated with various concentrations of the BTK inhibitors (0.1 µM, 0.5 µM, or 1 µM) or DMSO and then administered to VWF HA1 mice, followed by laser-induced thrombus formation. The BTK inhibitor-treated human platelets were fluorescently labeled and infused continuously through a catheter inserted into the femoral artery. Their behavior in response to laser-induced vascular injury was monitored in real time using two-channel confocal intravital microscopy (Furie and Furie, J. Clin. Invest.2005, 115(12), 2255-62). Upon induction of arteriole injury untreated platelets rapidly formed thrombi with an average thrombus size of 6,450 ± 292 mm ² (mean ± s.e.m.), as shown in FIG.67 and FIG.68. Similarly, Formula (2) (1 μM) treated platelets formed a slightly smaller but not significantly different thrombi with an average thrombus size of 5733 ± 393 mm ² (mean ± s.e.m.). In contrast, a dramatic reduction in thrombus size occurred in platelets pretreated with 1 μM of Formula (10) (ibrutinib), 2600 ± 246 mm ² (mean ± s.e.m.), resulting in a reduction in maximal thrombus size by about 61% compared with control (P > 0.001) (FIG.67 and FIG.69). Similar results were obtained with platelets pretreated with 500 nM of Formula (2) or ibrutinib: thrombus size of 5946 ± 283 mm ² , and 2710 ± 325 mm ² respectively. These initial results may provide some mechanic background and explanation on the reported 44% bleeding related adverse event rates in the Phase III

RESONATE study comparing ibrutinib with ofatumumab. The results obtained for Formula (17) (CC-292) were similar to that for Formula (10) (ibrutinib), as shown in FIG.67, 68, and 69. The effect of the BTK inhibitor concentration is shown in FIG.70. These results demonstrate the surprising advantage of the BTK inhibitor of Formula (2), which does not interfere with thrombus formation, while the BTK inhibitors of Formula (17) (CC-292) and Formula (10) (ibrutinib) interfere with thrombus formation.

[00747] The objective of this study was to evaluate in vivo thrombus formation in the presence of BTK inhibitors. In vivo testing of novel antiplatelet agents requires informative biomarkers. By utilizing a genetic modified mouse von Willebrand factor (VWFR1326H) model that supports human but not mouse platelet-mediated thrombosis, we evaluated the effects of Formula (2), Formula (17) (CC-292), and Formula (10) (ibrutinib) on thrombus formation. These results show that Formula (2) had no significant effect on human platelet-mediated thrombus formation while Formula (10) (ibrutinib) was able to limit this process, resulting in a reduction in maximal thrombus size by 61% compared with control. Formula (17) (CC-292) showed an effect similar to Formula (10) (ibrutinib). These results, which show reduced thrombus formation for ibrutinib at physiologically relevant concentrations, may provide some mechanistic background for the Grade≥ 3 bleeding events (e.g., subdural hematoma, gastrointestinal bleeding, hematuria and postprocedural hemorrhage) that have been reported in ≤ 6% of patients treated with Formula (10) (ibrutinib).

[00748] GPVI platelet aggregation was measured for Formula (2) and Formula (10) (ibrutinib). Blood was obtained from untreated humans, and platelets were purified from plasma-rich protein by centrifugation. Cells were resuspended to a final concentration of 350,000/μL in buffer containing 145 mmol/L NaCl, 10 mmol/L HEPES, 0.5 mmol/L Na ₂ HPO ₄ , 5 mmol/L KCl, 2 mmol/L MgCl ₂ , 1 mmol/L CaCl ₂ , and 0.1% glucose, at pH 7.4. Stock solutions of Convulxin (CVX) GPVI were prepared on the day of experimentation and added to platelet suspensions 5 minutes (37 °C, 1200 rpm) before the induction of aggregation. Aggregation was assessed with a Chronolog Lumi-Aggregometer (model 540 VS; Chronolog, Havertown, PA) and permitted to proceed for 6 minutes after the addition of agonist. The results are reported as maximum percent change in light transmittance from baseline with platelet buffer used as a reference. The results are shown in FIG.71.

[00749] In FIG.72, the results of CVX-induced (250 ng/mL) human platelet aggregation results before and 15 minutes after administration of the BTK inhibitors to 6 healthy individuals are shown.

[00750] The results depicted in FIG.71 and FIG.72 indicate that the BTK inhibitor of Formula (10) (ibrutinib) significantly inhibits GPVI platelet aggregation, while the BTK inhibitor of Formula (2) does not, further illustrating the surprising benefits of the latter compound.

Example 13– Synergistic Combinations of BTK Inhibitors and CD19 Inhibitors

[00751] Combination experiments may be performed to assess the synergistic behavior of drug combinations of BTK inhibitors and CD19 inhibitors. Syngeneic animal models for B cell lymphomas, including murine models, may be used to assess this behavior using intravenous, intrasplenic, intraperitoneal, subcutaneous, intramuscular, or intracerebral induction.

[00752] A mouse xenograft model may also be used to assess the synergistic behavior of drug combinations of BTK inhibitors and CD19 inhibitors In this study 6 to 10-week-old NOD- SCID-γc–/– (NSG) mice are obtained from the Jackson Laboratory (Bar Harbor, ME) and the CD19+ human ALL line Nalm-6 is obtained from American Type Culture Collection (ATCC; Manassas, VA). Animals are injected via the tail vein with 10 ⁶ viable Nalm-6 cells in 0.2 mL of sterile phosphate buffered saline (PBS). Starting at 3 to 5 weeks after injection, mice are screened for engraftment every 3 weeks by retro-orbital sampling of peripheral blood.

Engraftment is detected by flow cytometric analysis using antibodies recognizing CD19 and CD45. In addition, mice are screened using anti-CD3 and anti-CD33 to ensure engraftment was of the lymphoblast population and not of a T-cell or myeloid lineage. Xenografted mice were randomized to treatment with (i) the BTK inhibitor of Formula (2), (ii) a murine anti-CD19 monoclonal antibody (BioXcell, Clone 1D3), (iii) a combination of the BTK inhibitor of Formula (2) and the same murine anti-CD19 monoclonal antibody, or (iv) vehicle, after establishment of disease, defined as more than 5% blasts detected in peripheral blood. Response to treatment was measured weekly by retro-orbital bleeds to determine white blood counts and percent blasts by flow cytometry.

Example 14– In Vivo Study of CD19 CAR-T Cells and BTK Synergistic Combinations

[00753] To evaluate the effect of inhibiting BTK in tumors treated with CD19 CAR-T cells, eight-week-old SCID-Beige mice (Charles River) mice are injected by tail vein on day 1 with 1 × 10 ⁶ green fluorescent protein-firefly luciferase (GFP-FFLuc) transduced NALM-6 tumor cells. Seven days after tumor engraftment, mice are injected intravenously with 1 × 10 ⁷ CAR-T cells (day 7 and day 14). Tumor bioluminescence is monitored using a Xenogen in vivo Imaging System (IVIS; Caliper Life Sciences, Hopkinton, MA) for acquisition of imaging data sets. To evaluate a BTK inhibitor effect, GFP-FFLuc transduced NALM-6 cells are injected via the tail vein, and mice are treated with (A) oral Formula (2) at a dose of 15 mg/kg BID, (B) intravenous CAR-T cells as described above, or (C) a combination of Formula (2) at a dose of 15 mg/kg BID and intravenous CAR-T cells as described above. The CAR-T cells may use a variety of signaling domains, including CD28, 4-1BB, and CD3ζ domains, different transmembrane domains, and different CD19 scFv domains to evaluate the optimal combination.